Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
176. _Ascent of the Dorataspida._--The group #Cladophracta#, or those
ACANTHARIA whose lattice-shell arises by the union of transverse
processes of the twenty radial spines, includes five different
families, whose stem-group is the family Dorataspida, with a simple
spherical lattice-shell. This family itself is, however, diphyletic in
origin, being composed of two essentially and originally different
subfamilies--Diporaspida and Tessaraspida (p. 803). The Diporaspida (p.
808, Pls. 137, 138) have been developed from the Phractacanthida, and as
each radial spine of the latter bears two opposite apophyses, so the
lattice-shell of the former has forty primary aspinal pores (two on the
base of each spine). On the other hand, the Tessaraspida (p. 830, Pls. 135,
136) have been developed from the Stauracanthida, and as each radial spine
of the latter bears four crossed apophyses, so the lattice-shell of the
former has eighty primary aspinal pores (four at the base of each spine).
177. _Descent of the Diporaspida._--Whilst the Tessaraspida (§ 176) have
given rise to no new groups which could take rank as independent families,
no less than four separate families of ACANTHARIA have arisen from the
Diporaspida. The Phractopeltida (Pl. 133, figs. 1-6) are distinguished from
all other ACANTHARIA by the possession of two concentric spherical
lattice-shells, and have probably been developed from the {cxvi}Diporaspida
in the same way as the Dyosphærida from the Monosphærida among the
#Sphæroidea#; in that case the smaller inner lattice-sphere (medullary
shell) would be the primary, and the larger outer sphere (cortical shell)
the secondary; this latter shows forty primary aspinal pores like those of
the Diporaspida. The possibility is not excluded, however, that the small
inner lattice-sphere of the Phractopeltida is a secondary product. The
three remaining families, which must be regarded as descendants of the
Diporaspida, form together a single phylogenetic series, and are separated
from the primitive group mainly by the fact that the original spherical
form of the lattice-shell has been modified into one distinguished by an
elongated equatorial axis (the hydrotomical axis); hence the #Prunophracta#
(pp. 794-859). The ellipsoidal Belonaspida have arisen directly by
hypertrophy of the two opposite equatorial spines of this hydrotomical axis
(p. 859, Pl. 136, figs. 6-9; Pl. 139, figs. 8, 9; perhaps they have also
arisen directly from the Amphilonchida). In the lentelliptical Hexalaspida
(Pl. 139) all six spines which lie in the hydrotomical meridian plane (two
equatorial and four polar) are very strongly developed, the remaining
fourteen being rudimentary. Finally, in the Diploconida the two conical
sheaths of the two opposite hydrotomical equatorial spines are so
predominant that they take the chief part in the formation of the
hour-glass-shaped shell.
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