Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
186. _Genealogical Tree of the Plectoidea._--The order #Plectoidea#
includes those NASSELLARIA whose rudimentary skeleton does not contain the
characteristic sagittal ring of the #Stephoidea#, but consists of several
(at least three) radial spines, which proceed from a point in the centre of
the porochora. The branches of these radial spines remain free in the
Plagonida, whilst in the Plectanida they unite with each other to form a
loose meshwork (not, however, a complete lattice-shell). The number and
arrangement of the radial spines, which serve for generic distinctions, are
the same in both families, so that each genus of the Plectanida has arisen
from a corresponding genus of the Plagonida. The simplest Plagonida, which
possess a basal tripod (_Triplagia_ or _Plagiacantha_ with three rays,
_Tetraplagia_ with four rays) are probably to be regarded as forming the
common origin of the whole order. These agree with certain three- and
four-rayed skeletal pieces of the #Beloidea# (Thalassosphærida and
Sphærozoida); and also the four and six-rayed twinned pieces of the latter
(spicula bigemina and trigemina) repeat in the same fashion the skeleton of
the former (_Plagonidium_, _Plagonium_). This similarity, however, is a
mere analogy and possesses no phylogenetic significance. On the other hand,
certain Plagonida (_Plagoniscus_, _Plagiocarpa_), and the corresponding
genera of Plectanida (_Plectaniscus_, _Periplecta_) seem to have important
phylogenetic relations to certain #Stephoidea# (_Cortina_, _Cortiniscus_,
&c.); the sagittal ring of the latter having perhaps arisen by the vertical
apical spine of the former having been connected with their horizontal
basal rod by two ventral apophyses growing out opposite to each other
(compare pp. 902, 914, _Plagiocarpa procortina_, Pl. 91, fig. 5). In this
case the Plectanida would belong to the simplest stem-forms of the
NASSELLARIA.
187. _Genealogical Tree of the Stephoidea._--The order #Stephoidea#
includes all those NASSELLARIA whose skeleton does not form a complete
lattice-shell, but consists of one or more rings, and often of a loose
meshwork which arises by the union of branches of the rings. A _vertical
sagittal ring_ is constantly present, embracing the central capsule in the
median sagittal plane, and forming at its basal pole various processes, the
starting point for other skeletal forms. The most important of these is the
tripodal _Cortina_ {cxxi}(p. 950, § 182). The Stephanida are the most
archaic family among the #Stephoidea# (p. 937, Pl. 81), perhaps indeed
among all the NASSELLARIA (§ 184); in them the sagittal ring and its
processes alone constitute the skeleton; secondary rings and meshes are
wanting. Two diverging families, the Semantida and Coronida, have been
developed from the Stephanida, and from one of them the family Tympanida
has arisen.
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