Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The inception and digestion of nutriment, as it usually appears to take
place by the pseudopodia, has already been so fully treated in my
Monograph (L. N. 16, pp. 135-140), and since then in my paper on the
sarcode body of the Rhizopoda (L. N. 19, p. 342), that I have nothing of
importance to add. Quite recently Karl Brandt has expressed a doubt as to
whether the taking up of formed particles by the pseudopodia and their
aggregation in the calymma be really connected with the process of
nutrition. He is disposed rather to believe that these foreign bodies are
usually only accidentally and mechanically brought into the calymma, and
that the nourishment of the Radiolaria is derived exclusively or
pre-eminently from the symbiotic Xanthellæ (L. N. 52, pp. 88-93). I must,
however, maintain my former opinion, which I have only modified insomuch
that I now regard the sarcodictyum (on the outer surface of the calymma,
§ 94) rather than the sarcomatrix (on the outer surface of the central
capsule, § 92) as the principal seat of true digestion and assimilation.
From the sarcodictyum the dissolved and assimilated nutritive matters may
pass by the intracalymmar pseudopodia (or sarcoplegma, § 93) into the
sarcomatrix, and hence may reach the endoplasm through the openings in
the central capsule. To what an extent the Radiolaria are capable of
taking up even large formed bodies into the calymma, is shown by the
{cxxxi}striking instance of _Thalassicolla sanguinolenta_, which becomes
so deformed by the inception of numerous coccospheres and coccoliths,
that I described it as a special genus under the name _Myxobrachia_
(compare pp. 23, 30; also L. N. 21, p. 519, Taf. xviii., and L. N. 33, p.
37).
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