Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
C. On the active horizontal creeping movements of mature Radiolaria on a
firm ground, compare L. N. 12, p. 10, and L. N. 16, pp. 132-134.
D. Regarding the motion of social central capsules within the same
coenobium and the changes thus brought about in the structure of the
calymma, see L. N. 16, pp. 119-127, and L. N. 52, pp. 75-82.
221. _Contraction._--Motions, which are due to the contraction of
individual portions and cause changes in volume or form, have been partly
already spoken of under the head of locomotion (§ 220) and are partly
connected with other functions. Examples may be seen in the contraction of
the central capsule and of the calymma. A certain {cxlii}contraction of the
central capsule is probably brought about by the myophanes, which arise by
differentiation of the endoplasm and hence may assume different forms in
the four legions. In the SPUMELLARIA, where numerous radial fibrillæ run
from the central nucleus to the capsule membrane (§ 77), the endoplasm is
probably driven out evenly through all the pores of the capsule membrane by
their simultaneous contraction, and hence the volume of the capsule is
diminished in all directions. The ACANTHARIA probably behave similarly,
but are different, inasmuch as the number of their contractile radial
fibrillæ is less, and special axial threads (§ 78) are already
differentiated. In the NASSELLARIA it is probable that owing to the
contraction of the divergent myophane fibrillæ in the podoconus the
vertical axis of the latter is shortened, the opercular rods of the
porochora are lifted, and the endoplasm driven out of its pores, so that
the volume of the monaxon central capsule is diminished (§ 79). In the
PHÆODARIA the same result is probably brought about by the contraction of
the cortical myophane fibrillæ, which run meridionally along the inside of
the capsule membrane from the apical to the basal pole of the vertical main
axis, where they are inserted into the periphery of the astropyle; since
the volume of the capsule is diminished by their contraction (their
spheroidal figure becoming more nearly spherical) the endoplasm will be
driven out through the proboscis of the astropyle. Whilst these
contractions of the central capsule are largely due to differentiated
muscle-like threads of endoplasm (myophanes), this appears to be but rarely
the case with the contractions of the extracapsulum (_e.g._, the myophriscs
of the #Acanthometra#, § 96). Most of the phenomena of contraction which
can be observed in the calymma and pseudopodia depend upon exoplasmatic
currents (§ 209).
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