Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The new and interesting group, which was thus erected into an order under
the name TRIPYLEA, I had already a year previously separated from the other
Radiolaria as "_Pansolenia_" in my Protistenreich (L. N. 32, p. 102).
Since, however, neither the three capsular openings of the TRIPYLEA nor the
skeletal tubes of the Pansolenia are present in all the families of this
extensive order, I substituted in 1879 the more suitable name PHÆODARIA,
which is applicable to all members of the group (L. N. 34). In the
preliminary memoir then published regarding the Phæodaria, a New Group of
Siliceous Marine Rhizopods, I distinguished four orders, ten families, and
thirty-eight genera. The great majority of these new forms (among which
were no less than 465 different species) were first discovered by the
deep-sea investigations of the Challenger. John Murray was the first who
called attention to the great abundance in the deep sea of these remarkable
Rhizopods, and to the constant presence of their peculiar, dark,
extracapsular pigment body (phæodium); even in 1876 he described a portion
of them as Challengerida (L. N. 27, p. 536; L. N. 53, p. 226). The earliest
observations on the PHÆODARIA were made at Messina in 1859, where I
examined five genera of this remarkable group alive (compare p. 1522 and L.
N. 16).
By the discovery that the PHÆODARIA, although differing in important
respects from the other Radiolaria, still conform to the definition of the
class, a new and extensive series of forms was added to this latter, and by
their closer investigation a fresh source of interesting morphological
problems was disclosed. In other groups, however, morphology was advanced
by comparative anatomical studies. In addition to the smaller contributions
of various authors, mentioned in the foregoing bibliography, I may
specially refer to the valuable Beiträge zur Kenntniss der
Radiolarien-Skelete, insbesondere der der Cyrtida by O. Bütschli (L. N. 40,
1882). On the basis of careful comparative anatomical studies,
investigations into the skeletal structure of a number of fossil
#Cyrtoidea# and critical application of the recently published researches
of Ehrenberg into the Polycystina of Barbados (L. N. 25), Bütschli
attempted to derive the complicated relations of the Monopylean skeletons
phylogenetically from a simple primitive form,--the primary sagittal ring.
Even if this attempt did not actually solve the very difficult
morphological problem in question, still the critical and synthetic mode in
which it was carried out deserves full recognition, and furnishes the proof
that the comparative anatomy of the skeleton in the Radiolaria not less
than in the Vertebrata, is a most interesting and fruitful field of
phylogenetic investigation. A {clxxxii}further demonstration of this was
furnished by Bütschli in the general account of the organisation of the
Radiolaria which he published in 1882 in Bronn's Klassen und Ordnungen des
Thierreichs (L. N. 41).
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