Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
(_e.g._, in _Tuscaridium_). The parapylæ seem to be quite absent in the
families Challengerida, Medusettida, Castanellida, and perhaps also in
other PHÆODARIA. The form and structure of the small accessory openings
appear to be always the same. The {xxxii}outer capsule-membrane is elevated
in the form of a short cylindrical tube or "apertural ring" (collare
paraboscidis), the external margin of which bends inwards, and at the base
of the ring passes over into the delicate internal capsule membrane. Upon
this apertural ring is situated a longer or shorter "apertural cone"
(paraboscis), which is a tubular, cylindrical or conical, prolongation of
the membrane, open externally.
The peculiar capsule-openings of the PHÆODARIA were first discovered and
carefully described by Hertwig in 1879 (L. N. 33, pp. 95, 107). He found
in all the six genera which he examined _three_ openings, a main-opening
at the basal pole of the main axis and two accessory openings, one on
either side of the apical pole; hence he named the whole group
"TRIPYLEA." This name, however, is not applicable to the numerous
PHÆODARIA mentioned above, which have only a main opening without any
accessory openings, nor to those genera in which the number of the latter
is variable. I have, therefore, replaced Hertwig's designation by the
term "CANNOPYLEA," which has reference to the peculiar tubular form of
the opening. This I find much more developed in many PHÆODARIA than
Hertwig has represented, and I must also, in certain particulars, dissent
from his delineation of the minute structure, although this is in the
main remarkably accurate.
61. _The Nucleus._--The nucleus, enclosed in the central capsule of all
Radiolaria, behaves in every respect like a true cell-nucleus, and thus
lies at the base of the now universal opinion, that the whole Radiolarian
organism, in spite of its varied development and remarkable variations, is
unicellular and remains throughout life a true individual cell. This
important theory is not invalidated by the fact that the nucleus undergoes
peculiar modifications in many groups, and in certain groups presents
appearances seldom or never seen elsewhere.
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