62. _Uninuclear and Multinuclear Radiolaria (Monocaryotic and Polycaryotic)._--All Radiolaria present two different conditions in respect of the behaviour of the nucleus, since in their young stages they are uninuclear (_monocaryotic_), and in later stages multinuclear (_polycaryotic_). This is readily explained by the fact that each individual Radiolarian is developed from a simple unicellular swarm-spore, and that afterwards, before the formation of swarm-spores, the single nucleus divides into many small nuclei. Thus in the Radiolaria the nucleus is pre-eminently the _organ of reproduction and inheritance_. The division of the originally single nucleus into many small nuclei may take place, however, at very different periods, so that the Radiolaria may be divided in this respect into precocious and serotinous. 63. _Serotinous and Precocious Radiolaria._--In the great majority of the Radiolaria the division of the nucleus takes place only at a late period, a short time or even immediately before the process of spore formation; it then breaks up rapidly into numerous small nuclei (always more than one hundred, sometimes many thousands), and each of these {xxxiii}either becomes itself the nucleus of a swarm-spore, or by repeated division gives rise to a group of spore-nuclei. All those Radiolaria which are uninuclear during the greater part of their existence, and in which the process of division is late, and takes place rapidly, are called "serotinous" or late-dividing forms. To this category belong all PHÆODARIA and NASSELLARIA, as well as all the solitary or monozoic SPUMELLARIA and some ACANTHARIA. On the other hand, the name "precocious," or early dividing, is applied to those Radiolaria in which the division of the nucleus takes place very early, and in which, therefore, the cell is multinuclear during the greater part of its existence. This is the case in all the social or polyzootic Radiolaria (Polycyttaria, Pls. 3-8), and also in the great majority of the ACANTHARIA, both #Acanthometra# and #Acanthophracta#. In the last two groups, however, there are numerous exceptions, and these are seen in remarkably large species, characterised by the great size of the central capsule. From a phylogenetic point of view, the conclusion is allowable that the precocious forms are secondary, and have arisen by adaptive modification from the primitive serotinous stem. In the Polycyttaria (or social SPUMELLARIA, _i.e._, the three families Collozoida, Sphærozoida, and Collosphærida), the cause of the adaptation lies most probably in the formation of the colony itself, for all these three families are so closely related to three corresponding families of serotinous, monozootic Radiolaria (Thalassicollida, Thalassosphærida, Ethmosphærida), that certain species of the latter are hardly to be distinguished from isolated individuals of the former. Perhaps the remarkable formation of the large