Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
73. _The Intracapsular Fat-Globules._--Fat is present in the central
capsule of all Radiolaria in larger or smaller quantities, and generally
appears in the form of very {xliii}numerous, small, spherical granules,
which are either distributed evenly in the endoplasm (as an emulsion) or
enclosed in the vacuoles; the latter, in particular, is the case in most
PHÆODARIA, perhaps generally. In this group each vacuole contains as a rule
a single dark, shining fat-granule, and sometimes also an irregular bunch
composed of from two to five or more granules. In addition to these small
fat-granules (_granula adiposa_) which are always present, the central
capsule of many Radiolaria contains also larger fat-globules (_globuli
adiposi_). These appear to be generally wanting in the PHÆODARIA, and are
on the whole rare in the ACANTHARIA; whilst, on the contrary, they are very
common in the NASSELLARIA and SPUMELLARIA. The Polycyttaria or social
Radiolaria are as a rule distinguished by the possession of a single large
central oil-globule, which lies in the centre of the central capsule, and
is on an average about one-third of it in diameter (Pl. 3, figs. 4, 5).
This is absent, however, in those young capsules of the Polycyttaria in
which the primary nucleus is centrally situated (Pl. 3, fig. 12). Those
species of Polycyttaria whose central capsule reaches a considerable size,
often enclose numerous oil-globules, and in _Collophidium_ (species of
_Collozoum_ with an elongated cylindrical capsule, Pl. 3, figs. 1, 3) the
axis of each capsule is occupied by a row of numerous oil-globules. In the
monozootic SPUMELLARIA, in which the nucleus is always centrally situated,
the large oil-globules are, of course, excentric, being in apposition to
the inner surface of the capsule-membrane (Pl. 1, fig. 3; Pl. 2, figs. 2,
5). In the #Discoidea# the oil-globules, which are often present in large
numbers, form elegant concentric rings around the central nucleus, and in
those species with segmented arms, there are one or more transverse rows in
each segment (Pl. 43, fig. 15). In the NASSELLARIA the number and
distribution of the oil-globules are dependent upon the form of the central
capsule. When this is simple, without lobes, and ovoid or conical, they
generally lie in its aboral half above the podoconus (Pl. 51, figs. 5, 13;
Pl. 97, fig. 1). When, on the contrary, the basal portion of the capsule
sends out three or four dependent processes (as in the majority of the
#Cyrtoidea#), a large globule may generally be seen in the swollen distal
part of each conical or ovoid lobe (Pl. 53, fig. 19; Pl. 60, figs. 4-7). In
many #Stephoidea# and #Spyroidea#, whose central capsule is separated into
two lateral portions by the constriction corresponding to the sagittal
ring, each of these contains either a single large globule or a group of
small ones (Pl. 90, figs. 7, 10). These oil-globules are usually colourless
and highly refractive; rarely they are yellow or brown, sometimes
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