Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
A. The fine fibrillæ in the cortical layer of the endoplasm were first
described by Hertwig in 1879 (L. N. 33, p. 98, Taf x. figs. 6-10). He
found them, however, only below the three openings in the capsule of the
Tripylea, where they form three stellate groups of fibrils. I find them
very clearly shown, and with especial distinctness, under the astropyle
in most PHÆODARIA of which I have had the opportunity of examining
well-stained and preserved central capsules. In many cases, also, the
striation is not confined to the apertures, but spreads over the whole
cortical layer. Perhaps this constitutes in all PHÆODARIA a thin
myophane-sheet, whose contractile fibrils run from one pole of the main
axis to the other and cause by their contraction changes in the form of
the spheroidal central capsule.
B. The granular medullary portion of the endoplasm of the PHÆODARIA, with
its numerous clear spherical vacuoles, was first described in my
Monograph (1862), in the case of _Aulacantha_ (p. 263), _Aulosphæra_ (p.
359), and _Coelodendrum_ (p. 361) as a "finely granular, mucous substance
(intracapsular sarcode), packed more or less closely with clear spherical
vesicles from 0.005 to 0.015 mm. in diameter, each of which contains one
or two, rarely three, dark shining granules." That these clear spheres
are true vacuoles was first clearly proved by Hertwig (L. N. 33, p. 98).
As a rule all the vacuoles of the same central capsule are of equal size
(generally from 0.008 to 0.012 mm. in diameter), and are distributed at
equal intervals throughout the finely granular endoplasm.
{li}CHAPTER III.--THE EXTRACAPSULUM.
(§§ 81-100).
81. _The Components of the Extracapsulum._--The extracapsulum or
extracapsular malacoma, under which name are included all those parts of
the soft body which lie outside the central capsule, consists of the
following constant, and important constituents:--(1) The _calymma_ or
extracapsular jelly-veil; (2) the _sarcomatrix_ or layer of exoplasm
immediately surrounding the membrane of the central capsule; (3) the
_sarcodictyum_ or network of exoplasm, covering the surface of the calymma;
(4) the _pseudopodia_ or radial fibres of exoplasm, which may again be
subdivided into intracalymmar pseudopodia, uniting the sarcomatrix and
sarcodictyum, and extracalymmar pseudopodia, radiating freely into the
water outside the calymma.
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