Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
imbibition (L. N. 16, pp. 109, 110). R. Hertwig was the first to
demonstrate, in 1879, that the jelly-veil is constantly present in living
Radiolaria, that it forms the basis of the extracapsular malacoma and
surrounds the central capsule as a second protective sheath (L. N. 33, p.
114).
83. _The Structure of the Calymma._--The extracapsular jelly-veil appears
structureless in most Radiolaria, inasmuch as it represents a homogeneous
pellucid excretion of the exoplasm and contains neither fibres nor other
formed structures. In some groups, however, definite structural characters
become secondarily developed. The most common and striking of these is the
formation of alveoles, which takes place in the extracapsulum (see § 86).
In consequence of this the calymma assumes a remarkable frothy consistency
and appears to be composed of large, clear, thin-walled vesicles; this is
especially the case in the #Collodaria# (#Colloidea#, Pls. 1, 3, and
#Beloidea#, Pls. 2, 4), and in many large PHÆODARIA, especially among the
#Phæocystina# (Phæodinida and Cannorrhaphida, Pl. 101, and Aulacanthida,
Pls. 102-104). More rarely the calymma is not permeated by vacuoles, but
there appear in it fine striæ parallel to the surface as though it were
composed of thin concentric laminæ like an onion; perhaps these are the
expressions of a different quantity of water in the various layers. In the
calymma of many Radiolaria thin, straight, radial lines are to be seen,
which are probably pseudopodia, and not to be attributed to any structural
modification, or they may be slender canals which serve for the exit of the
pseudopodia. On the outer surface of the calymma of different Radiolaria,
and especially in the ACANTHARIA, a peculiar network of fibres is to be
found, composed of polygonal meshes, like elastic fibres, probably due to a
local thickening of the jelly. These polygonal meshes are often very
regularly distributed between the radial spines of the #Acanthometra#, and
stand in a definite relation to them. The fibres which form the meshes are
often rather strong, resembling elastic fibres, as above-mentioned, and
either simple or composed of bundles of very fine fibrillæ (L. N. 33, p.
15, Taf. i. fig. 1, Taf. ii. fig. 4).
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