Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
91. _The Exoplasm or Extracapsular Protoplasm._--The extracapsular
protoplasm, which may be shortly termed the "exoplasm" (or ectosarc), is
primitively in all Radiolaria (and especially in their earliest development
stages) the only important constituent of the extracapsulum, besides the
calymma. Although the extracapsular and intracapsular protoplasm of the
Radiolaria are everywhere in direct communication, and although the
openings in the membrane of the central capsule bring about an interchange
between them, still the two portions of sarcode show certain constant and
characteristic differences, which are due to the physiological division of
labour between the central and peripheral parts of the body and their
corresponding morphological differentiation. The extracapsular, like the
intracapsular, protoplasm is originally homogeneous, but may afterwards
become differentiated in various ways, producing the special constituents
of the extracapsulum. Such "external protoplasmic products" are vacuoles,
pigment-bodies, &c. More important, however, are the topographically
different sections into which the exoplasm may be divided according to its
relations to the central capsule and the calymma. In this respect the
following parts may be generally distinguished--(1) the _Sarcomatrix_, or
fundamental layer of the exoplasm, which surrounds the central capsule as a
continuous sheath of sarcode and separates it from the calymma; (2) the
_Sarcoplegma_, an irregular network of the exoplasm, which spreads
throughout the gelatinous material of the calymma; (3) the _Sarcodictyum_
or network of sarcode on the outer surface of the calymma; and (4) the
_Pseudopodia_, which project outwards from the latter and radiate into the
water.
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