Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
lattice-shell whatever, the conformation of the sarcodictyum is usually
irregular, with meshes of irregular form and unequal size; sometimes,
however, they seem to be very regular, as in many #Acanthometra# (Pl. 129,
fig. 4).
95. _The Pseudopodia._--On the whole the pseudopodia or thread-like
processes of the exoplasm exhibit in the Radiolaria the same characteristic
peculiarities as in all true Rhizopoda; they are usually very numerous,
long and thin, flexible and sensitive filaments of sarcode, which show the
peculiar phenomena of granular movement. Their physiological significance
is in several respects very great, for they serve as active organs for the
inception of nutriment, for locomotion, sensation, and the formation of the
skeleton (see note A, below). The presence of a calymma, however, which
distinguishes the Radiolaria from the other Rhizopoda, brings about certain
modifications in the behaviour of the pseudopodia. If in general all the
threads, which arise from the sarcomatrix or fundamental layer and radiate
outwards, be called "pseudopodia," then that part of them which is included
in the gelatinous substance of the calymma and forms the sarcoplegma may be
termed the "collopodia" (or intracalymmar pseudopodia), and the remaining
portion, which passes outwards from the sarcodictyum freely into the water,
may be described as "astropodia" (or extracalymmar pseudopodia). In many
Radiolaria these two portions present some differences in morphological and
physiological respects, and certain distinctions are probably generally
present (see note B). Apart from this universal differentiation in the
different groups of the Radiolaria, specially modified forms of pseudopodia
may be recognised as the axopodia and myxopodia of the ACANTHARIA (see §
95, A), and the sarcode-flagellum of certain SPUMELLARIA (see note C).
A. The pseudopodia of the Radiolaria have been so fully described in my
Monograph, in 1862, both morphologically and physiologically, that I need
only refer to the account there given {lxiii}(pp. 89-127); for
supplementary observations see R. Hertwig (1879, L. N. 33, p. 117) and
Bütschli (1882, L. N. 41, pp. 437-445).
B. The _Astropodia_, or free radiating pseudopodia, are in many
Radiolaria more or less clearly distinguishable from the collopodia,
which form the sarcoplegma within the calymma; how far these distinctions
depend upon a permanent differentiation (especially in the ACANTHARIA and
PHÆODARIA) needs further investigation.
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