Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
On more violent irritation and after the death
of the #Acanthometra# the myophriscs separate from the radial bars and
remain attached to the distal ends of the conical gelatinous sheaths as
free "ciliary coronas." At the same time, {lxv}they melt into short, thick,
hyaline rods, the so-called "gelatinous cilia." The myophriscs are found
only in the order #Acanthometra#, and are wanting in the #Acanthophracta#,
as well as in the other three legions of Radiolaria.
A. The "_ciliary coronas_" on the skeletal rods of dead #Acanthometra#
were first described by the discoverer of this order, Johannes Müller,
and referred to as "the stumps of the contracted, thickened threads" (L.
N. 12, p. 11, Taf. xi.).
B. The "_number of the gelatinous cilia_" I found constant in certain
species of #Acanthometra#, and stated in my Monograph (L. N. 16, p. 115)
"that here is to be found the first differentiation of the diffuse
sarcode into definite organs of regular definite number, size, and
position, which deserve the name tentacles rather than pseudopodia."
C. The nature of the myophriscs as fibrillæ allied to muscles was first
discovered by R. Hertwig, who described them as "structures of peculiar
nature," under the name of "contractile threads," and pointed out in
detail their histological and physiological peculiarities (L. N. 33, pp.
16-19, Taf. i.).
97. _The Exoplasm of the Peripylea._--The extracapsular protoplasm of the
SPUMELLARIA or PERIPYLEA is in communication with the intracapsular sarcode
by the innumerable fine pores of the capsule-membrane, and like these pores
is evenly distributed over the whole surface. The sarcomatrix which
immediately surrounds the central capsule is moderately strong, and sends
out innumerable long, thin pseudopodia, which probably correspond to the
pores of the membrane. Their number is markedly greater in the SPUMELLARIA
than in the other three legions. The ramifications and communications which
the radiating fibres of the sarcomatrix undergo within the calymma,
apparently present the most manifold variations, so that the sarcoplegma or
intracalymmar network thus formed has very diverse forms. On the surface of
the calymma the exoplasmic threads constitute a variously disposed
sarcodictyum, a regular or irregular exoplasmic network, by the
silicification of which a primary lattice-shell arises in the majority of
the SPUMELLARIA. The free ends of the pseudopodia, which arise from this
extracalymmar network and radiate out into the water, appear in most
SPUMELLARIA to be relatively short, but exceedingly numerous. Specially
modified pseudopodia and axial threads in particular do not seem to occur
in this legion. Perhaps, however, among the latter may be reckoned the
remarkable pseudopodia which combine to form the sarcode flagellum in many
#Discoidea# (and perhaps in other SPUMELLARIA). This axoflagellum is a
particularly strong thread of sarcode, arising from a definite point in the
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