Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia): Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
100. _The Exoplasm of the Cannopylea._--The extracapsular protoplasm of the
PHÆODARIA or CANNOPYLEA is much better developed as regards volume than in
the other three legions, and is connected with the intracapsular sarcode by
only a few apertures in the capsule-membrane. In most PHÆODARIA three of
these are present, the astropyle or main-opening at the oral pole of the
main axis, and the two lateral parapylæ or accessory openings on either
side of the aboral pole (§ 60). In several families the latter appear to be
wanting, whilst in others their number is increased; these families have
not yet, however, been observed during life. The protoplasm projects both
from the oral main-opening and from the two aboral accessory openings in
the form of a thick cylindrical rod; the tube into which each opening is
produced in many PHÆODARIA (longer in the case of the astropyle, shorter in
the parapylæ) being regarded as an excretion from this protoplasmic
cylinder. The sarcode threads within the tube appear like a bundle of
fibrils, either quite hyaline or finely striated. After issuing from the
mouth of the aperture they pass over into a thick sarcomatrix, which
surrounds the central capsule entirely and separates it from the enclosing
calymma. In the neighbourhood of the basal astropyle the sarcomatrix is
usually swollen into a thick lenticular disc, which is in direct contact
with the peculiar phæodium of this legion (§ 89). The pseudopodia, which
radiate from the sarcomatrix, and form by anastomosis a wide-meshed
sarcoplegma within the calymma, are usually not very numerous in the
PHÆODARIA, but are very strong. Sometimes two stronger bundles of
collopodia may be distinguished at the two poles of the main axis, an oral
bundle (in the direction of the proboscis of the astropyle) and an aboral
bundle (at the opposite pole between the parapylæ). The collopodia of the
sarcoplegma unite at the surface of the calymma into a regular or irregular
sarcodictyum, which, in most PHÆODARIA produces by the secretion of a
peculiar silicate the primary lattice-shell. {lxviii}The free astropodia,
which pass outwards from the sarcodictyum into the water, are in most
PHÆODARIA very numerous (Pl. 101, fig. 10). Since, however, only a few
species of this great legion have been observed in a living state, their
pseudopodia require further accurate examination.
CHAPTER IV.--THE SKELETON.
(§§ 101-140).
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