Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The thorax of the #Botryodea#, or the second shell-joint, absent in the
Cannobotryida, is usually of very simple form, ovate, truncate, conical or
cylindrical, and consists of a secondary joint, developed from the base of
the cephalis. The thorax is usually small, about the same size as the
cephalis or a little larger, whilst in the #Cyrtoidea# it is commonly much
larger than the latter. Its terminal mouth is closed by a lattice-plate in
the Botryocellida, while it remains open in the Botryopylida and
Pylobotryida.
The abdomen, or the third shell-joint, developed in the Pylobotryida only,
generally also exhibits a very simple shape, like the thorax, and it is a
tertiary joint developed from the base of the thorax. It exhibits to the
latter the same relation as in the Tricyrtida. The terminal mouth of the
abdomen remains open in the Botryocyrtida; it becomes closed by a
lattice-plate in the Botryocampida.
The lattice-plate forming the shell of the #Botryodea# is usually very thin
and fragile, with very small circular pores. These are often very numerous,
at other times scarce, and sometimes nearly wanting, so that the shell
appears hyaline. In this group there never occurs that great variety in
form and size of the pores, which numerous #Cyrtoidea# and #Spyroidea#
exhibit.
The radial apophyses which are found in the majority of #Botryodea# seem to
correspond in position and relation to the typical radial beams of the
other NASSELLARIA, viz., three descending basal feet arising from the base
of the cephalis, and an ascending vertical apical horn arising from the top
of the cephalis. Here also they seem to be inherited from the #Plectoidea#
or #Stephoidea# (_Cortina_, _Cortiniscus_). But whilst these four typical
radial beams in the other NASSELLARIA are usually simple spines or solid
rods, here they usually appear as hollow cylindrical tubes, the thin wall
of which is pierced by the same small pores as the shell. These porous
tubes are either straight or slightly curved, often inflated and ovate at
the base. We find in their number and disposition the same variety as in
the simple radial spines of the other NASSELLARIA. The original number may
here also be seen in the development of four typical beams, an ascending
apical tube on the top of the cephalis, and three descending basal tubes on
its base; the odd posterior tube of the latter corresponds to the caudal
foot, the two paired anterior to the pectoral feet. There is often also a
fourth foot developed opposite to the caudal, and representing a sternal
tube. Sometimes also two superior tubes are developed, a posterior
occipital and an anterior frontal. The different {1107}apophyses may be
lost by reduction. I have never observed #Botryodea# with more than five
apophyses.
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