Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The thorax, or the second shell-joint of the #Cyrtoidea#, is in the
majority homologous with the thorax of the Phormospyrida and Androspyrida,
and therefore developed by apophyses, which arise from the base of the
cephalis and become united by transverse branches forming a lattice-plate.
Its size is generally in inverse proportion to that of the cephalis. The
more the cephalis becomes reduced, the more the thorax is developed. Its
form is very variable, usually three-sided pyramidal or prismatic in the
triradiate, polyhedral in the multiradiate, and conical or cylindrical in
the eradiate #Cyrtoidea#. Its terminal mouth is either a simple wide
opening, or closed by a lattice-plate. In the majority of #Cyrtoidea# the
thorax is separated from the cephalis not only by the external collar
constriction, but also by the internal cortinar septum, a horizontal
lattice-plate which exhibits the typical basal pores of the Semantida
(usually two smaller jugular and two larger cardinal pores). But this
septum is often reduced or perfectly lost, and then the external collar
constriction alone indicates the separation of the cephalis and the thorax.
The abdomen, or the third shell-joint of the #Cyrtoidea#, absent in the
Monocyrtida and Dicyrtida (as also in all #Spyroidea#), occurs constantly
in all Tricyrtida and Stichocyrtida. It is a simple large chamber in the
Tricyrtida, but forms an annulated body, composed of a variable number of
successive joints, in the Stichocyrtida. The constrictions between these
joints, and also the lumbar constriction, between abdomen and thorax, are
usually provided with a lattice-girdle, projecting into the cavity of the
shell, like a diaphragm. Usually this horizontal girdle bears only a single
circle of pores, rarely two or more. In many #Cyrtoidea# it is replaced by
a solid horizontal ring of silex, and often it is wanting. It originates by
the insertion of the following shell-joint, which takes place not on the
terminal mouth of the preceding joint, but somewhat above it.
{1131}The annular joints of the Stichocyrtida succeeding the third joint,
and very variable in number, may be regarded either as a series of new
postabdominal chambers, succeeding the true abdomen, or as secondary joints
of the annulated abdomen itself. The latter view may be sustained by the
fact that these joints are usually of an indifferent shape, and do not
possess the characteristic features which we find in the first three
joints, the abdomen, the thorax and the cephalis.
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