Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
{1193}The family #Tripocyrtida#, composed of the Sethopilida and
Sethoperida of my Prodromus, comprises those #Cyrtoidea# in which the
lattice-shell is two-jointed and bears three radial apophyses. The two
subfamilies differ in the shape of the mouth, which in the Sethopilida is a
simple wide opening, but in the Sethoperida is closed by a lattice-plate;
the former are here divided into sixteen, the latter into eight different
genera. Though probably the two shell-joints are not truly homologous in
all Tripocyrtida, we call the first joint here, as in all Dicyrtida, the
cephalis, and the second joint the thorax.
Numerous Tripocyrtida, living as well as fossil forms, were formerly
described by Ehrenberg. His genera _Dictyophimus_, _Clathrocanium_,
_Lithomelissa_, and _Lychnocanium_ belong to the Sethopilida, and have the
mouth open; his genera _Lithopera_ and _Lithochytris_ (partly) belong to
the Sethoperida, and have the basal mouth closed by a lattice-plate. Many
of these Tripocyrtida belong probably to the oldest forms of Dicyrtida, are
nearly related to the Phormospyrida, and therefore of special phylogenetic
interest, as was demonstrated by Bütschli (1882, _loc. cit._, pp. 514-519).
This near relation to certain #Spyroidea# (_Tripospyris_, _Acrospyris_,
&c.) is particularly striking in some forms of _Clathrocanium_,
_Lithomelissa_, &c. Some other Tripocyrtida seem to possess a closer
relation to certain #Plectoidea# (_Plagoniscus_, _Plectaniscus_), so mainly
some forms of _Tripocyrtis_ and _Dictyophimus_.
The cephalis, or the first joint of the shell, corresponds usually to the
whole shell of the Zygospyrida and of numerous Monocyrtida, and exhibits
various modifications of shape, which have been already described in these
latter families. It is usually subspherical or hemispherical and armed with
an apical horn. In a small number of genera the horn is lost, in some other
genera multiplied. The cephalis is separated from the thorax not only
externally by the collar constriction, but commonly also internally by a
transverse horizontal fenestrated septum, which usually exhibits three or
four characteristic cortinar pores. The central capsule, originally
enclosed in the cephalis, develops usually three or four large pear-shaped
cæcal sacs which pass through the cortinar pores and depend into the thorax
(Pl. 55, figs. 2-11; Pl. 60, figs. 3-7, &c.).
The thorax in this family exhibits a great variety of interesting
modifications, mainly in the development of the three radial apophyses
arising from it. These may be either enclosed in the wall of the thorax as
ribs, or arise as free wings, very often prolonged over the mouth as three
terminal feet. Finally the three terminal feet only remain, whilst the
original ribs are lost. The special ornamentation of these three apophyses
exhibits an extraordinary variety and elegancy of structure, and many
Tripocyrtida belong, no doubt, to the most graceful and admirable forms of
NASSELLARIA.
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