Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=166. Plant types.= The necessity for decreasing or increasing water
loss in compensation of the water supply has made it possible to
distinguish two fundamental groups of plants upon the twofold basis of
habitat and structure. These familiar groups, xerophytes and
hydrophytes, represent two extremes of habitat and structure, between
which lies a more or less vague, intermediate condition represented by
mesophytes. These show no characteristic modifications, and it is
consequently impossible to arrange them in subgroups. Xerophytes and
hydrophytes, on the other hand, exhibit marked diversity among
themselves, a fact that makes it desirable to recognize subgroups, which
correspond to fundamental differences of habitat or adaptation. It is
hardly necessary to point out that these types are not sharply defined,
or that a single plastic species may be so modified as to exhibit
several of them. The extremes are always clearly defined, however, and
they indicate the specific tendency of the adaptation shown by other
members of the same group.
=167. Xerophytic types.= With the exception of dissophytes, all
xerophytes agree in the possession of a deep-seated root system, adapted
to withdraw water from the lower moist layers, and to conserve from loss
from the upper dry layers. Reservoirs are developed in the root,
however, in relatively few cases. The stem follows the leaf more or less
closely in its modification, except when the leaf is greatly reduced or
disappears, in which event the stem exhibits peculiar adaptations. While
the leaf is by far the most strikingly modified, it is a difficult task
to employ it satisfactorily as the basis for distinguishing types.
Several adaptations are often combined in the same leaf, and it is only
where one of these is preeminently developed, as in the case of
succulence, that the plant can be referred to a definite type. The
latter does not happen in many species of the less intensely xerophytic
habitats, and, consequently, it is difficult, if not undesirable, to
place such xerophytes under a particular group. The best that can be
done is to recognize the types arising from extreme or characteristic
modification, and to connect the less marked forms as closely as
possible with these. Halophytes differ from xerophytes only in the fact
that the chresard is determined by the salt-content of the habitat, and
not by the texture of the soil. In consequence, they should not be
treated as a distinct group.
[Illustration: Fig. 33. Staurophyll of _Bahia dissecta_, showing extreme
development of palisade (chresard, 3–9%; light, 1). × 130.]
=168. Types of leaf xerophytes.= In these, adaptation has acted
primarily upon the leaf, while the stem has remained normal for the most
part. Even when the leaves have become scale-like, they persist
throughout the growing season, and continue to play the primary part in
photosynthesis. The following types may be distinguished:
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account