Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
2. _The floating type._ With respect to form and the structure of the
upper part of the leaf, floating leaves are essentially similar to those
of amphibious plants. They are usually lacquered or coated with wax to
prevent the stoppage of the stomata by water. Stomata, except as
vestiges, are found only on the upper surface, and the palisade tissue
is much less developed than the sponge, which is uniformly characterized
by large air-chambers. The stems are elongated, the aerating system is
enormously developed, and the supportive tissues are reduced. In the
_Lemnaceae_, the leaf and the stem are represented by a mere frond or
thallus, and the roots are in the process of disappearance, e. g.,
_Spirodela_ has several, _Lemna_ one, and _Wolffia_ none.
3. _The submerged type._ Both stem and root have been greatly reduced in
submerged plants, owing to the generalization of absorption and the
density of the water. The leaves are greatly reduced in size and
thickness, chiefly, it would seem, for the purpose of insuring readier
aeration and great illumination. The leaf may be ribbon-like, linear,
cylindrical, or finely dissected. Stomata are sometimes present, but
they are functionless and vestigial. A distinction into palisade and
sponge tissues, when present, must also be regarded as a vestige; the
chlorenchym is essentially that of a shade leaf. The air-chambers are
much reduced, and sometimes lacking; they function doubtless as
reservoirs for air obtained from the water.
PHOTOHARMOSE
_ADJUSTMENT_
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