Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=286. Influence of levels.= The invasion of a formation may occur at
three different levels: (1) at the level of the facies, (2) below the
facies, (3) above the facies, depending directly upon the relative
height of invaders and occupants. The invasion level is an extremely
simple matter to determine, except in the case of woody plants, such as
shrubs and trees, which attain their average height only after many
years. Its importance is fundamental. The level at which invasion occurs
not only determines the immediate constitution of the formation, whether
its impress shall still be given by the occupants or by the invaders or
by both together, but it also decides the whole future of the formation,
i. e., whether the invaders or occupants shall persist unmodified or
modified. The problem is an extremely complex one, but the careful
analysis of invasion at each level throws a flood of light upon it. The
entrance of invaders of the same general height as the facies of a
formation results regularly in mixed formations. This is well
illustrated by the structure of the transition areas between two
formations of the same category, i. e., forests, meadows, etc. It is
seldom, however, that the facies and invaders are so equally matched in
height and other qualities that they remain in equilibrium for a long
period. One or the other has a slight advantage in height, or the one
suffers shading or crowding better than the other, is longer-lived or
faster-growing, with the result that invader yields to occupant, or
occupant to invader. It is a well-known fact that many mixed formations
represent intermediate stages of development.
Invasion at a level different from that of the facies is inevitably
followed by modification. If the invasion takes place below the facies,
the invaders will be exterminated gradually, or slowly assimilated. In
either case, there is little structural change in the formation, and its
stability is affected slightly or not at all. If the invaders overtop
the facies in any considerable number, the entire formation undergoes
partial or complete modification, or in extreme cases it disappears, as
is typically the case in succession. A peculiar variation of invasion at
a level above the facies is seen where woody plants invade grassland,
when the trees or shrubs become more or less uniformly scattered in an
open woodland or open thicket. Here the grassland takes on an altogether
different appearance superficially, though it is usually unchanged,
except beneath and about the invaders, where either adaptation or
extermination results. Finally, it should be borne in mind that the
invasion of a particular formation, especially in the case of layered
thickets and forests, often takes place at two levels, at the height of
the facies and below the facies.
_INVESTIGATION OF INVASION_
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