Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
the case of bad lands, or, more frequently, they are mesotropic, passing
first through a long series of xerophytic formations. Sandbars
(_cheradia_, _syrtidia_) should be considered here, though they are
eroded by currents and waves, and not by run-off. They are fixed and
built up by sand-binding grasses and sedges, usually of a hydrophytic
nature, and pass ultimately into mesophytic forest.
=319. Succession by filling with silt and plant remains.= All aquatic
habitats into which silt, wash, or other detritus is borne by streams,
currents, floods, waves, or tides are slowly shallowed by the action of
the water plants present. These not only check the movement of the
water, thus greatly decreasing its carrying power, and causing the
deposition of a part or all of its load, but they also retain and fix
the particles deposited. In accordance with the rule, each plant becomes
the center of a stabilizing area, which rises faster than the rest of
the floor, producing the well-known hummocks of lagoons and swamps. All
aquatics produce this reaction. It is more pronounced in submerged and
amphibious forms than in floating ones, and it takes place more rapidly
with greatly branched or dissected plants than with others. In pools
(_tiphia_) and lakes (_limnia_), debouching streams and surface waters
deposit their loads in consequence of the check exerted by the still
water and the marginal vegetation, and delta-like marshes are quickly
built up by filling. Springs (_crenia_) likewise form marshes where they
gush forth in sands, the removal of which is impeded by vegetation. The
flood plains and deltas of rivers show a similar reaction. The heavily
laden flood waters are checked by the vegetation of meadows and marshes,
and deposit most of their load. The banks of streams (_ochthia_) and of
ditches (_taphria_) are often built up in the same fashion by the action
of the marginal vegetation upon the current. The presence of marginal
vegetation often determines the checking or deflecting of the current in
such a way as to initiate meanders, while natural levees owe their
origin to it, in part at least. Along low seacoasts, waves and tides
hasten the deposit of river-borne detritus, causing the water to spread
over the lowlands and form swamps. They often throw back also the
sediment that has been deposited in the sea, the marsh vegetation acting
as a filter in both cases. Successions of the kind indicated above are
regularly mesotropic. Where the soil is sandy, and the filling-up
process sufficiently great, or where salts or humus occur in excess,
xerophytic formations result. In certain cases, these successions appear
to be permanently hydrostatic, changing merely from floating or
submerged to amphibious conditions, but this is probably due to the
slowness of the reaction. As a rule, the accumulation of plant remains
is relatively slight, and plays an unimportant part in the reaction. In
peat bogs and other extensive swamps, the amount of organic matter is
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