Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
(5) Excessive seed production and slight mobility lead to the
imperfect ecesis of individuals in dense stands, and in
consequence usually produce great instability.
(6) Each pioneer produces about itself a tiny area of ecesis and
stabilization for its own offspring, for the disseminules of
its fellows, or of invaders.
(7) Species propagating by offshoots, or producing relatively
immobile disseminules in small number, usually show effective
ecesis, as the offspring appear within the area of the reaction
of the parent forms.
V. Stabilization.
(1) Stabilization is the universal tendency of vegetation.
(2) The ultimate stage of a succession is determined by the
dominant vegetation of the region. Lichen formations are often
ultimate in polar and niveal zones; grassland is the final
vegetation for plains and alpine stretches, and for much
prairie, while forest is the last stage for mesophytic midlands
and lowlands, as well as for subalpine regions.
(3) Grassland or forest is the usual terminus of a succession;
they predominate in lands physiographically mature.
(4) The limit of a succession is determined in large part by the
progressive increase in occupation, which makes the entrance of
invaders more and more difficult.
(5) Stabilization proceeds radiately from the pioneer plants or
masses. The movement of offshoots is away from the parent mass,
and the chances of ecesis are greatest near its edges, in a
narrow area in which the reaction is still felt, and the
occupation is not exclusive.
VI. General laws.
(1) The stages, or formations, of a succession are distinguished
as initial (_prodophytia_), intermediate (_ptenophytia_), and
ultimate (_aiphytia_).
(2) Initial formations are open, ultimate formations are closed.
(3) The number of species is small in the initial stages; it
attains a maximum in intermediate stages; and again decreases
in the ultimate formation, on account of the dominance of a few
species.
(4) The normal sequence of vegetation forms in succession is: (1)
algae, fungi, mosses; (2) annuals and biennials; (3) perennial
herbs; (4) bushes and shrubs; (5) trees.
(5) The number of species and of individuals in each stage
increases constantly up to a maximum, after which it gradually
decreases before the forms of the next stage. The interval
between two maxima is occupied by a mixed formation.
(6) A secondary succession does not begin with the initial stage
of the primary one which it replaces, but usually at a much
later stage.
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