Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
The slope of a habitat exerts a considerable effect upon the intensity
of the incident light. If the angle between the slope and the sun’s ray
be 90°, a square meter of surface will receive the maximum intensity, 1.
At an angle of 10°, the same area receives but .17 of the light. This
relation between angle and intensity is shown in the table which
follows. The influence of the light, however, is felt by the leaf, not
by the slope. Since there is no connection between the position of the
leaf and the slope of the habitat, the latter may be ignored. In
consequence, it is unnecessary to make allowances for the direction of a
slope, viz., whether north, east, south, or west, in so far as light
values are concerned. The angle which a leaf makes with its stem
determines the angle of incidence, and hence the amount of light
received by the leaf surface. This is relatively unimportant for two
reasons. This angle changes hourly and daily with the altitude of the
sun, and the intensity constantly swings from one extreme to the other.
Moreover, the extremes 1.00 and 0.17, even if constant, are hardly
sufficient to produce a measurable result. When the angle of the leaf
approaches 90°, there is the well-known differentiation of leaf surfaces
and of chlorenchym, but this has no relation to the angle of incidence.
_Table of Intensity at Various Angles_
ANGLE INTENSITY
90 1.00
80 .98
70 .94
60 .87
50 .77
40 .64
30 .50
20 .34
10 .17
In the sunlight, it makes no difference at what height a light reading
is taken. In forest and thicket as well as in some herbaceous
formations, the intensity of the light, if there is any difference, is
greatest just beneath the foliage of the facies. In forests especially,
the light is increasingly diffuse toward the ground, particularly where
layers intervene. In woodland formations, moreover, the exact spot in
which a reading is made must be carefully chosen, unless the foliage is
so dense that the shade is uniform. A very satisfactory plan is to take
readings in two or more spots where the shade appears to be typical, and
to make a check reading in a “sunfleck,” a spot where sunlight shows
through. In forests and thickets, the sunflecks are fleeting, and the
light value is practically that of the shade. In passing into open
woodland and thicket, the sunflecks increase in size and permanence,
until finally they exceed the shade areas in amount and become typical
of the formation.
_Reflected and Absorbed Light_
Public-domain text, read in full here on John Shaqi.
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