Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
Maculæ figured of Phac. (Acaste) Downingiæ. SALTER Monogr. pl. 2 fig.
34 b, p. 25 »high up on each side a small tubercle».
_Phac. rana_ HALL Pal. N. Y. vol. VII, pl. VIII A, fig. 18. Hypostoma
with the large commalike maculæ high up.
Of the following species Ph. cephalotes and Ph. macrophthalmus belong
to the genus Phacops proper, while Phac. quadrilineata, as well as
Ph. Downingiæ are of a quite different generic type. The name Acaste
proposed for them is not good as LEACH in 1817 named a Cirrhiped as
Acasta.
=Ph. cephalotes= BARR.
Pl. V figs. 35, 36.
The hypostoma is elongate, almost triangular, tricuspidate at the
posterior border, a little above this there is a crescent-shaped ridge,
and still a little higher above this and the median horizontal line,
the two macula, as diminutive tubercles. Seen with magnifying powers
they are slightly oblong or reniform.
=Ph. macrophthalmus= BURM.
Pl. VI figs. 1-9.
The triangular hypostoma has a little below the median horizontal
diameter two shallow grooves, and a little above the same diameter the
extraordinary minute maculæ may with some care be observed, figs. 8,
9. They are visible as a small bare, smooth spot amongst the curious
elevated and incoherent terrace lines of the surface. The exceedingly
small size of these maculæ naturally does not allow any research of
their structure. They are rather in this species and many of the
congeneric to be regarded as small ocelli. In some specimens these
spots are quite rudimentary and are reduced to the smallest area
possible. The anterior wings are of a peculiar earlike shape, finely
striate, enlarged and flat near the hypostoma and ending in a little
hollow horn (figs. 6, 7). A little below it, the posterior wing emerges
as a small angular knob. The aggregate cephalic eyes are more distinct.
As the vertical section learn us the regular ovate lenses consist
generally of clear, crystalline calcareous spar and are on the surface
covered by an extremely thin membrane, not a common one for all, but
a separate cornea for each lens, which envelopes its superior moiety
and continues down as a thickened appendix between the lens and the
interstitial test (figs. 3-5). In the horizontal sections it surrounds
the lenses as an annular wall. Around some lenses there is as if still
another ring lay outside this, but more indistinct, probably only
clearer portions of the test surrounding them. The cheeks close to the
lenses are indistinctly perforated by longitudinal tubes.
=Phacops (Acaste) quadrilineata= ANGELIN.
Pl. V figs. 37, 38.
We have not succeeded in finding any well preserved hypostoma.
The lenses of the cephalic eyes resemble much those of the preceding
species though more elongate and ovate. They are covered by a membrane
which surrounds their upper moiety and between the lenses joins the
interstitial test. In the horizontal section these membranes surround
the lenses as an annular wall.
=Phaëtonides= BARR.
=Ph. Stokesi= MURCH.
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