Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
32 | Phillipsia | 5 | -- |
33 | Platymetopus | 1 | -- |
34 | Pliomera | 1 | -- |
35 | Proetus | 3 | -- |
36 | Ptychoparia | 1 | -- |
37 | Ptychopyge | 4 | 2 |
38 | Schmidtia | 1 | -- |
39 | Trochurus | 2 | -- |
| +===========+=====================+
| Total | 136 | 36 |
[Footnote 42: To these have here been added two Bohemian species P.
bohemicus and P. spinosus omitted above at page 64.]
We have thus 136 species of 39 genera in which hypostomic maculæ have
been found and only 36 species, in which it has been possible to study
the structure of the maculæ through sections.
Common for a great number of maculæ in various groups, whether they
show any organic structure or not, is the excessive thinness of their
shell in comparison with that of the surrounding hypostoma. This is
also in accordance with the tenuity of the cephalic eyes in relation to
the test of the cheeks.
The structure which characterizes the macula as a visual organ,
although often so rudimentary, is not in all instances spread through
the whole substance of the macula. This structure has in several
species been restricted to a narrow circumference of the body of the
macula, to its interior apex as in Bronteus and Proetus. Only in the
Asaphidæ, in Illænus and Lichas the entire macula shows this structure.
Perhaps, to judge by certain indications in Bronteus, once in a larval
or preceding stage of evolution the whole surface of the macula was
also in that genus covered with lenses, which have been reduced.
The different groups in which the examined genera may be divided are
the following seven, showing the remarkably great diversity of these
organs. There is even in the same genus so great a variability that
species with structure in the macula occur along with those devoid of
any structure or also, as in Lichas, with a different structure. It
must, however, be remembered, that the species of such genera may not
be coeval.
_Group 1._ In the sectioned macula there is no trace whatever of any
structure. The test of the macula considerably thinner than that
of the hypostoma. According to the affinities of the genera they
may be subdivided as follows. _a_ Bumastus, Dysplanus. _b_ Nileus,
Symphysurus, ? Ogygia. _c_ Calymmene, Homalonotus. _d_ Chirurus pro
parte.
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