Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
The position of the hypostomic eyes on the ventral surface of the
trilobites is not quite so abnormous, nor so isolated a feature amongst
the crustaceans as might at first be supposed. Amongst the recent
crustacea there is probably none, which as to the conformation of its
labrum or hypostoma resembles the trilobites so closely as the species
of the genus Apus, however different they may else be in all other
respects. We have studied chiefly the hypostoma of Apus cancriformis.
In its shape as seen from the outside and as to its outlines it reminds
strongly of the trilobite hypostoma. It is nearly square (pl. VI figs.
46, 47) with rounded corners. The anterior margin is arched and in
the middle it has a broad tonguelike prolongation. On the sides of
the anterior margin there are two short, sharply pointed wings. The
lateral margins are thick, almost as doubled, and bent forward so that
a groove is formed alongside them and the large, evenly vaulted median
surface of the hypostoma. These grooves are continued in a short groove
parallel with the posterior margin. From this margin a narrow oblique
surface is slanting backwards, as it were, doubling that margin. On
its surface there are two small, resplendent, white or clear spots,
almost of the shape of a crescent, though more irregular when seen
in higher enlargement (pl. VI fig. 49). They are translucent as to
be visible on the interior side of the hypostoma. In Apus glacialis
there are also similar spots in the same position, but they form short
narrow, straight stripes, parallel with the border of the posterior
margin. I have no opinion as to the nature or function of these white
spots. I do not think that they at all are homologous with the macula;
of the trilobites. On the other hand we see a little higher up on the
exterior side of the hypostoma two faintly elevated elliptic tubercles
which as to their position and shape highly remind of the maculæ of the
trilobites. They are hollow on the inside and we have not been able to
find any peculiar structure in them. That they have any visual function
is precluded through the circumstance that the whole exterior surface
is densely covered with microscopic bristles (pl. VI fig. 49) which are
especially remarkable along the posterior margin.
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