Response in the Living and Non-LivingBose, Jagadis Chandra
Science
Response in the Living and Non-Living
Bose, Jagadis Chandra
Electrophysiology; Fatigue; Irritability
Just as in the strain cells the galvanometer contact was transferred
from the electrolytic part to the metallic part of the circuit, so we
may next, in an exactly similar manner, cut this plate into two, and
connect these directly to the galvanometer, electrolytic connection
being made by partially plunging them into a cell containing water. The
posterior surfaces of the two half-plates may be covered with a
non-conducting coating. And we arrive at a typical photo-electric cell
(fig. 98, _b_). These considerations will show that the eye is
practically a photo-electric cell.
[Illustration: FIG. 99.--RESPONSES TO LIGHT IN FROG'S RETINA
Illumination L for one minute, recovery in dark for two minutes during
obscurity D. (Waller.)]
We shall now give detailed experimental results obtained with the
sensitive silver-bromide cell, and compare its response-curve with those
of the retina. A series of uniform light stimuli gives rise to uniform
responses, which show very little sign of fatigue. How similar these
response-curves are to those of the retina will be seen from a pair of
records given below, where fig. 99 shows responses of frog's retina, and
fig. 100 gives the responses obtained with the sensitive silver cell
(fig. 100).
It was said that the responses of the retina are uniform. This is only
approximately true. In addition to numerous cases of uniform responses,
Waller finds instances of 'staircase' increase, and its opposite, slight
fatigue. In the record here given of the silver cell, the staircase
effect is seen at the beginning, and followed by slight fatigue. I have
other records where for a very long time the responses are perfectly
uniform, there being no sign of fatigue.
[Illustration: FIG. 100.--RESPONSES IN SENSITIVE SILVER CELL
Illumination for one minute and obscurity for one minute. Thick line
represents record during illumination, dotted line recovery during
obscurity.]
Another curious phenomenon sometimes observed in the response of retina
is an occasional slight increase of response immediately on the
cessation of light, after which there is the final recovery. An
indication of this is seen in the second and fourth curves in fig. 99.
Curiously enough, this abnormality is also occasionally met with in the
responses of the silver cell, as seen in the first two curves of
fig. 100. Other instances will be given later.
CHAPTER XVIII
INORGANIC RESPONSE--INFLUENCE OF VARIOUS CONDITIONS ON THE RESPONSE TO
STIMULUS OF LIGHT
Effect of temperature--Effect of increasing length of exposure--Relation
between intensity of light and magnitude of
response--After-oscillation--Abnormal effects: (1) preliminary
negative twitch; (2) reversal of response; (3) transient positive
twitch on cessation of light; (4) decline and reversal--Resume.
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