In Echinoderms as a class, the body presents three regions differing
in structure, and on the greater or less development of these regions
or systems, as we may call them, their chief differences are based.
Take, for instance, the dorsal system, the nature of which is
explained by the name, indicating of course the back of the animal,
though it does not necessarily imply the upper side of the body, since
some of the Echinoderms, as the stemmed Crinoids, for example, carry
the dorsal side downward, while the Star-fishes and Sea-urchins carry
it upward, and the Holothurians, moving with the mouth forward, have
the dorsal system at the opposite end of the body. Whatever the
natural attitude of the animal, however, and the consequent position
of the dorsal region, it exists alike in all the five orders, though
it has not the same extent and importance in each. But in all it is
made up of similar parts, bears the same relation to the rest of the
body, has the same share in the general economy of the animal. And
though when we compare the spreading back of a Star-fish with the
small area on the top of a Sea-urchin, where all the zones unite, we
may not at once see the correspondence between them, yet a careful
comparison of all their structural details shows that they are both
built with the same elements and represent the same region, though it
is stretched to the utmost in the one case, and greatly contracted in
the other.
This being true of the dorsal system, let us look at another equally
important structural feature in this class. All Echinoderms have
locomotive organs peculiar to themselves, a kind of suckers which may
be more or less numerous, larger or smaller, in different species, but
are always appendages of the same character. These are variously
distributed over the body, but always with a certain regularity
occupying definite spaces, shown by investigation to be homologous in
all. For instance, the rays of the Star-fish correspond in every
detail on their under side, along which the locomotive suckers run,
with the zones on the Sea-urchin, from end to end of which the suckers
are arranged; and the same is equally true of the distribution of the
suckers on the Holothurians, Ophiurans, and Crinoids, though, as most
persons are less familiar with these orders than with the other two,
it might not be so easy to point out the coincidence to our readers.
These suckers are called the ambulacra, the lines along which they run
are called the ambulacral rows or zones, while the system of
locomotion as a whole is known as the ambulacral system. Since these
organs are thus regularly distributed over the body in distinct zones
or rows, it follows that the latter must be divided by intervening
spaces. These intervals are called the interambulacral spaces; but
while in some orders they are occupied by larger plates and prominent
spines, as in the Sea-urchin and Star-fish, in others they are either
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