The highest of our Holothurians in structure, is the Pentacta. (Fig.
130) It is very rare on our beaches, though occasionally found under
stones at low-water mark; farther north, in Maine, and at Grand Manan,
it is very common, covering all the rocks near low-water mark. It is a
chocolate brown in color, and measures, when fully expanded, some
fifteen to eighteen inches in length. Unlike the Cuvieria, the
ambulacral suckers are evenly distributed and almost equally developed
on all the tubes; between the five rows of ambulacral suckers are
scattered irregularly certain appendages resembling suckers, but found
on examination not to be true locomotive suckers, and called on that
account false ambulacra. These are the organs corresponding to the
warts around the mouth of the Synapta. Although the ambulacral suckers
are, as we have said, equally developed on all the tubes, yet the
Pentacta does not use them indiscriminately as locomotive organs. In
Pentacta, as well as in all Holothurians, whether provided with
ambulacral suckers, or, like the Synapta and Caudina, deprived of
them, the odd ambulacrum, viz. the one placed opposite the madreporic
body, is always used to creep upon, and forms the under surface of the
animal.
[Illustration: Fig. 130. Pentacta frondosa; expanded about one
third the natural size.]
The correspondence between the different phases of growth in the young
Pentacta, and the adult forms of the orders described above, the
Synapta, Caudina, Cuvieria, and Pentacta itself, is a striking
instance of the way in which embryonic forms illustrate the relative
standing of adult animals. In the earlier stages of its development,
the ambulacral tubes alone are developed in the Pentacta; in this
condition it recalls the lower orders of Holothurians, as the Synapta
and Caudina; then a sole is formed by the greater development of three
of the ambulacra, and in this state it reminds us of the next in
order, the Cuvieria, while it is only in assuming its adult form that
the Pentacta develops its other ambulacra, with their many suckers.
The Pentacta resembles the Trepang, so highly valued by the Chinese as
an article of food, and forms a not unsavory dish, having somewhat the
flavor of lobster.
* * * * *
ECHINOIDS.
_Sea-urchin_. (_Toxopneustes drobachiensis_ AG.)
[Illustration: Fig. 131. Toxopneustes from above, with all the
appendages expanded; natural size.]
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