The position of the pedicellariæ is quite different from that which
they occupy in the Sea-urchin, where they are scattered singly between
the spines and tentacles, though more regularly and closely grouped
along the lines upon which the refuse food is moved off. In the
Star-fish, on the contrary, these singular organs seem to be grouped
for some special purpose around the spines, on the upper surface of
the body. Every such spine swells near its point of attachment, thus
forming a spreading base (Fig. 143), around which the pedicellariæ are
arranged in a close wreath, in the centre of which the summit of the
spine projects; they differ also from those of the Sea-urchin in
having two prongs instead of three. Other pedicellariæ are scattered
independently over the surface of the animal, but they are smaller
than those forming the clusters and connected with the spines. The
function of these organs in the Star-fish remains unexplained; the
opening on the upper surface, through which the refuse food is thrown
out, is in such a position that they evidently do not serve here the
same purpose which renders them so useful to the Sea-urchin.
Occasionally they may be seen to catch small prey with these forks,
little Crustacea, for instance; but this is probably not their only
office. The Star-fish has a fourth set of external appendages in the
shape of little water-tubes. (Seen in Fig. 143.) The upper surface of
the back consists of a strong limestone network (Fig. 144), and
certain openings in this network are covered with a thin membrane
through which these water-tubes project. It is supposed that water may
be introduced into the body through these tubes; but while there can
be no doubt that they are constantly filled with water, and are
therefore directly connected with the circulation through the
madreporic body (Fig. 145), no external opening has as yet been
detected in them. The fact, however, that when these animals are taken
out of their native element, the water pours out of them all over the
surface of the back, so that they at once collapse and lose entirely
their fulness of outline, seems to show that water does issue from
those tubes. The ends of the arms are always slightly turned up, and
at the summit of each is a red eye-speck. The tentacles about the eye
become very delicate and are destitute of suckers.
[Illustration: Fig. 143. Single spine of Star-fish, with
surrounding appendages; magnified.]
[Illustration: Fig. 144. Limestone network of back of Star-fish.]
[Illustration: Fig. 145. Madreporic body of Star-fish; magnified.]
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