The red grouse is placed in a genus called "Lagopus," of which there
are several species in the northern hemisphere. In Scotland the red
grouse, which is distinguished as Lagopus Scoticus, is accompanied by
a rarer species of Lagopus, which lives in high, bare regions. This is
the bird called by the Celtic name "ptarmigan"; it differs in several
points from the red grouse, and acquires white plumage in the winter,
which the latter bird does not; it is called Lagopus mutus. Now in
Norway we find also two species of grouse or Lagopus, called "rypé"
(pronounced "reeper") by the Norwegians. One is the same bird in every
respect as the Scotch ptarmigan, and is known as "the mountain rypé."
The other is very close to our red grouse, and is called "the common
or bush rypé," and by English naturalists the "willow grouse," and
by ornithologists "Lagopus salicetus." It agrees in habits, voice,
eggs, and anatomical detail with our red grouse, but the back of the
cock-bird of the red grouse and the whole plumage of the hen-bird
have a darker colour. Moreover, the willow grouse, like the ptarmigan
or mountain rypé, turns white–acquires a white plumage–in the winter
which the red grouse does not. Are the red grouse and the willow grouse
to be regarded as distinct species? Our British red grouse lives on
heather-grown moors; the willow grouse prefers the shrubby growths of
berry-bearing plants interspersed with willows, whence its name. Their
food differs accordingly. Formerly the red grouse lived on the moors
of the South of England, and when in Pleistocene times England was a
part of the Continent of Europe the willow grouse and the red grouse
were one undivided species inhabiting all the north-west of Europe. It
is probable, though the experiment would be almost impossible to carry
out, that were the eggs of a number of willow grouse now brought to
Scotland and hatched on the moors, they would tend to keep apart from
the native red grouse, and not inter-breed with them, in which case we
should say that the Scotch form is a "species on the make," or, even,
a completed and distinct species. On the other hand, it is possible
that the two forms would freely pair with another, and that the colour
and winter coat of the one (probably that of the Scotch form if the
experiment were tried in Scotland) would predominate, and after some
generations no trace of the other strain would be observable.
CHAPTER X
SOME SPECIFIC CHARACTERS
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