The appendage thus alluded to by Sir EVERARD HOME is the grand "cul-de-sac," noticed by the Académic des Sciences, and the "division particulière," figured by CAMPER. It is of sufficient dimensions to contain ten gallons of water, and by means of the valve above alluded to, it can be shut off from the chamber devoted to the process of digestion. Professor OWEN is probably the first who, not from an autopsy, but from the mere inspection of the drawings of CAMPER and HOME, ventured to assert (in lectures hitherto unpublished), that the uses of this section of the elephant's stomach may be analogous to those ascertained to belong to a somewhat similar arrangement in the stomach of the camel, one cavity of which is exclusively employed as a reservoir for water, and performs no function the preparation of food.[1] [Footnote 1: A similar arrangement, with some modifications, has more recently been found in the llama of the Andes, which, like the camel, is used as a beast of burden in the Cordilleras of Chili and Peru; but both these and the camel are _ruminants_, whilst the elephants belongs to the Pachydermata.] [Illustration] Whilst Professor OWEN was advancing this conjecture, another comparative anatomist, from the examination of another portion of the structure of the elephant, was led to a somewhat similar conclusion. Dr. HARRISON of Dublin had, in 1847, an opportunity of dissecting the body of an elephant which had suddenly died; and in the course of his examination of the thoracic viscera, he observed that an unusually close connection existed between the trachea and oesophagus, which he found to depend on a muscle unnoticed by any previous anatomist, connecting the back of the former with the forepart of the latter, along which the fibres descend and can be distinctly traced to the cardiac orifice of the stomach. Imperfectly acquainted with the habits and functions of the elephant in a state of nature, Dr. HARRISON found it difficult to pronounce as to the use of this very peculiar structure; but looking to the intimate connection between the mechanism concerned in the functions of respiration and deglutition, and seeing that the proboscis served in a double capacity as an instrument of voice and an organ for the prehension of food, he ventured (apparently without adverting to the abnormal form of the stomach) to express the opinion that this muscle, viewing its attachment to the trachea, might either have some influence in raising the diaphragm, and thereby assisting in expiration, "_or that it might raise the cardiac orifice of the stomach, and so aid this organ to regurgitate a portion of its contents into the oesophagus_."[1] [Footnote 1: _Proceed. Roy. Irish Acad_., vol. iv. p. 133.]