Slowly the superb erection composed of the four flat fan-like pinions
assumes rigidity and colour. By to-morrow the colour will have attained
the requisite shade. For the first time the wings close fan-wise and lie
down in their places; the elytra bend over at their outer edges, forming
a flange which lies snugly over the flanks. The transformation is
complete. Now the great locust has only to harden its tissues a little
longer and to tan the grey of its costume in the ecstasy of the
sunshine. Let us leave it to its happiness, and return to an earlier
moment.
The four stumps which emerge from their coverings shortly after the
rupture of the corselet along its median line contain, as we have seen,
the wings and elytra with their innumerable nervures. If not perfect,
at least the general plan is complete, with all its innumerable details.
To expand these miserable bundles and convert them into an ample set of
sails it is enough that the organism, acting like a force-pump, should
force into the channels already prepared a stream of humours kept in
reserve for this moment and this purpose, the most laborious of the
whole process. As the capillary channels are prepared in advance a
slight injection of fluid is sufficient to cause expansion.
But what were these four bundles of tissue while still enclosed in their
sheaths? Are the wing-sheaths and the triangular winglets of the larva
the moulds whose folds, wrinkles, and sinuosities form their contents in
their own image, and so weave the network of the future wings and
wing-covers?
Were they really moulds we might for a moment be satisfied. We might
tell ourselves: It is quite a simple matter that the thing moulded
should conform to the cavity of the mould. But the simplicity is only
apparent, for the mould in its turn must somewhere derive the requisite
and inextricable complexity. We need not go so far back; we should only
be in darkness. Let us keep to the observable facts.
I examine with a magnifying-glass one of the triangular coat-tails of a
larva on the point of transformation. I see a bundle of moderately
strong nervures radiating fan-wise. I see other nervures in the
intervals, pale and very fine. Finally, still more delicate, and running
transversely, a number of very short nervures complete the pattern.
Certainly this resembles a rough sketch of the future wing-case; but
how different from the mature structure! The disposition of the
radiating nervures, the skeleton of the structure, is not at all the
same; the network formed by the cross-nervures gives no idea whatever of
the complex final arrangement. The rudimentary is succeeded by the
infinitely complex; the clumsy by the infinitely perfect, and the same
is true of the sheath of the wing and the final condition of its
contents, the perfect wing.
Public-domain text, read in full here on John Shaqi.
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