Larger burrowing animals also assist in the task of mixing uniformly the surface soils, and aiding root-penetration, as well as, in many cases, the conservation of moisture. Seton (loc. cit.) even claims that the pocket gophers (Thomomys) in a great degree replace the activity of the earthworms in the arid region, where they, together with the voles (commonly known there as field mice), exist in great numbers. Of course the work of these animals, as well as that of the prairie dogs, ground squirrels, badgers, etc., is incompatible with cultivation. But the effects of their burrows on the native vegetation, and the indications they give of the nature of the subsoil, are eminently useful to the land-seeker. Thus in the rolling sediment-lands of the Great Bend of the Columbia, the observer is surprised to see the “giant rye grass,” usually at home in the moist lowlands, growing preferably on the crests of the ridges bordering the horizon. Examination shows that this is due to the burrowing of badgers, whereby the roots of the grass are enabled to reach moisture at all times, even in that extremely arid region. CHAPTER X. SOIL AND SUBSOIL (_Continued_) THEIR RELATIONS TO VEGETATION. _Physical Effects of the Percolation of Surface Waters._—The muddy water formed by the beating of rains on the soil surface will, in penetrating the soil, carry with it the diffused colloidal clay to a certain depth into the subsoil. We should therefore expect that as a rule every subsoil will be more clayey than its surface soil; and this is found to be almost universally the case in the humid region. Subsoils are therefore almost always less precious and more retentive of moisture, as well as of plant-food substances in solution, than their surface soils, unless these are very rich in humus; and as the finest particles are usually those richest in available plant-food, it follows that subsoils will as a rule be found to contain larger supplies of the latter than the surface soil. Common experience as well as comparative analysis confirm both of these inferences so thoroughly, that it becomes unnecessary to adduce examples in this place. On the other hand, the reverse, upward movement of moisture caused by surface evaporation tends constantly to bring any soluble salts contained in the soil mass nearer to the surface, thus increasing the stock of easily available plant-food in the surface soil. In extreme cases, especially in the arid region, this accumulation of salts may become excessive, and seriously injurious to plant growth. (See “Alkali Soils”, chapters 21, 22.)