The terms variation and variety are used frequently when speaking of
hybrids. By crossing forms, which are already variable in the sense just
mentioned, it is easy to multiply the number of the types, and even in
crossing pure forms the different characters may be combined in
different ways, the resulting combinations [190] yielding new, and very
often, valuable varieties. But it is manifest that this form of
variation is of quite another nature from the variations of pure races.
Many hybrid varieties are quite constant, and remain true to their type
if no further crosses are made; many others are artificially propagated
only in a vegetative way, and for this reason are always found true.
Hybrid varieties as a rule were formerly confused with pure varieties,
and in many instances our knowledge as to their origin is quite
insufficient for sharp distinctions. To every student of nature it is
obvious, that crossing and pure variability are wholly distinct groups
of phenomena, which should never be treated under the same head, or
under the same name.
Leaving aside polymorphy, we may now discuss those cases of variability,
in which the changes themselves, and not only their final results play a
part. Of such changes two types exist. First, the ever-recurring
variability, never absent in any large group of individuals, and
determining the differences which are always to be seen between parents
and their children, or between the children themselves. This type is
commonly called "individual variability" and since this term also has
still other meanings, it has of late become customary to use instead the
term "fluctuating variability." [191] And to avoid the repetition of the
latter word it is called "fluctuation." In contrast to these
fluctuations are the so-called sports or single varieties, not rarely
denominated spontaneous variations, and for which I propose to use the
term "mutations." They are of very rare occurrence and are to be
considered as sudden and definite steps.
Lastly, we have to consider those varieties, which vary in a much wider
range than the ordinary ones, and seem to fluctuate between two opposite
extremes, as for instance variegated leaves, cultivated varieties with
variegated or striped flowers, double flowers and some other anomalies.
They are eversporting and ever-returning from one type to the other. If
however, we take the group of these extremes and their intermediates as
a whole, this group remains constant during the succeeding generations.
Here we find once more an instance of the seemingly contradictory
combination of high variability and absolute constancy. It means that
the range of variability has quite definite limits, which in the common
course of things, are never transgressed.
We may infer therefore that the word variability has such a wide range
of meanings that it ought never be used without explanation. [192]
Nothing indeed, is more variable than the signification of the term
variable itself.
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