Many other instances could be given. Smooth varieties are seldom
absolutely so, and if scattering hairs are found on the leaves or only
on some more or less concealed parts, they correspond in their character
to those of the species. So it is with prickles, and even the thornless
thorn-apple has fruits with surfaces far from smooth. The thornless
horse-chestnut [235] has in some instances such evident protuberances on
the valves of its fruits, that it may seem doubtful whether it is a pure
and stable variety.
Systematic latency may betray itself in different ways, either by normal
systematic marks, or by atavism. With the latter I shall deal at length
on another occasion, and therefore I will give here only one very clear
and beautiful example. It is afforded by the common red clover.
Obviously the clovers, with their three leaflets in each leaf, stand in
the midst of the great family of papilionaceous plants, the leaves of
which are generally pinnate. Systematic affinity suggests that the
"three leaved" forms must have been derived from pinnate ancestors,
evidently by the reduction of the number of the leaflets. In some
species of clover the middle of the three is more or less stalked, as is
ordinarily the case in pinnate leaves; in others it is as sessile as are
its neighbors. In a subsequent chapter I will describe a very fine
variety, which sometimes occurs in the wild state and may easily be
isolated and cultivated. It is an ordinary red clover with five leaflets
instead of three, and with this number varying between three and seven,
instead of being nearly wholly stable as in the common form. It produces
from time to time pinnate leaves, [236] very few indeed, and only
rarely, but then often two or three or even more on the same individual.
Intermediate stages are not wanting, but are of no consequence here. The
pinnate leaves obviously constitute a reversion to some prototype, to
some ancestor with ordinary papilionaceous leaves. They give proof of
the presence of the common character of the family, concealed here in a
latent state. Any other explanation of this curious anomaly would
evidently be artificial. On the other hand nothing is really known about
the ancestors of clover, and the whole conception rests only on the
prevailing views of the systematic relationships in this family. But, as
I have already said, further proof must be left for a subsequent
occasion.
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