Hitherto we have dealt with negative characters and tried to prove that
the conception of latency of the opposite positive characteristics is a
more natural explanation of the phenomenon than the idea of a complete
loss. We have now to consider the positive varieties, and to show that
it is quite improbable that here the species have struck out for
themselves a wholly new character. In some instances such may have been
the case, but then I should prefer to treat these rather as elementary
species. But in the main we will have to assume the latency of the
character in the species and its reassumption by the variety when
originating, as the most probable explanation.
Great stress is laid upon this conception by the fact, that positive
varieties are so excessively rare when compared with the common
occurrence [239] of negative ones. Indeed, if we put aside the radiate
and the color-varieties of flowers and foliage, hardly any cases can be
cited. We have dealt with this question in a former lecture, and may now
limit ourselves to the positive color-varieties.
The latency of the faculty of producing the red pigment in leaves must
obviously be accepted for nearly the whole vegetable kingdom. Oaks and
elms, the beautiful climbing species of Ampelopsis, many conifers, as
for instance _Cryptomeria japonica_, some brambles, the Guelder-rose
(_Viburnum Opulus_) and many other trees and shrubs assume a more or
less bright red color in the fall. During summer this tendency must have
been dormant, and that this is so, is shown by the young leaves of oaks
and others, which, when unfolding in the spring show a similar but paler
hue. Moreover, there is a way of awakening the concealed powers at any
time. We have only to inflict small wounds on the leaves, or to cut
through the nerves or to injure them by a slight bruising, and the
leaves frequently respond with an intense reddening of the living
tissues around and especially above the wounds. _Azolla caroliniana_, a
minute mosslike floating plant allied to the ferns, responds to light
and cold with a reddish tinge, and to shade or warmth with a pure green.
The foliage [240] of many other plants behaves likewise, as also do
apples and peaches on the insolated sides of the fruits. It is quite
impossible to state these groups of facts in a more simple way than by
the statement that the tendency to become red is almost generally
present, though latent in leaves and stems, and that it comes into
activity whenever a stimulus provokes it.
Public-domain text, read in full here on John Shaqi.
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