The conception of latency of characters as the common source of the
origination of varieties, either in the positive or in the negative way,
leads to some rules on variability, which are known under the names
given to them by Darwin. They are the rules of repeated, homologous,
parallel and analogous variability. Each of them is quite general, and
may be recognized in instances from the most widely distant families.
Each of them is quite evident and easily understood on the principle of
latency.
By the term of repeated variability is meant the well-known phenomenon,
that the same variety has sprung at different times and in different
[243] countries from the same species. The repetition obviously
indicates a common internal cause. The white varieties of blue- and
red-flowered plants occur in the wild state so often, and in most of the
instances in so few individuals that a common pedigree is absolutely
improbable. In horticulture this tendency is widely and vexatiously
known, since the repetition of an old variety does not bring any
advantage to the breeder. The old name of "conquests," given by the
breeders of hyacinths, tulips and other flower-bulbs to any novelty, in
disregard of the common occurrence of repetitions, is an indication of
the same experience in the repeated appearance of certain varieties.
The rule of parallel variations demands that the same character
occasionally makes its appearance in the several varieties or races,
descended from the same species, and even in widely distinct species.
This is a rule, which is very important for the general conception of
the meaning of the term variety as contrasted with elementary species.
For the recurrence of the same deviation always impresses us as a
varietal mark. Laciniated leaves are perhaps the most beautiful
instance, since they occur in so many trees and shrubs, as the walnut
tree, the beech, the birch, the hazelnut, and even in [244] brambles and
some garden-varieties of the turnip (_Brassica_).
In such cases of parallel variations the single instances obviously
follow the same rules and are therefore to be designated as analogous.
Pitchers or ascidia, formed by the union of the margins of a leaf, are
perhaps the best proof. They were classified by Morren under two heads,
according to their formation from one or more leaves. Monophyllous
pitchers obey the same law, viz.: that the upper side of the leaf has
become the inner side of the pitcher. Only one exception to this rule is
known to me. It is afforded by the pitchers of the banyan or holy
fig-tree, _Ficus religiosus_, but it does not seem to belong to the same
class as other pitchers, since as far as it has been possible to
ascertain the facts, these pitchers are not formed by a few leaves as in
all other cases, but by all the leaves of the tree.
Public-domain text, read in full here on John Shaqi.
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