But recalling our former consideration of this subject the supposition
seems allowable that the tendency to revert is not connected with the
type of the hybrid, but is apt to occur in some rare individuals of
every type. But since most of the sterile hybrids are only known to us
in a single individual and its vegetative offspring, this surmise offers
an explanation of the rare occurrence of sports.
Finally, we must consider some of the so called hybrid races or strains
of garden-plants. _Dahlia_, _Gladiolus_, _Amaryllis_, _Fuchsia_,
_Pelargonium_ and many other common flowers afford the best known
instances. Immeasurable variability seems here to be the result of
crossing. But on a closer inspection the range of characters is not so
very much wider in these hybrid races than in the groups of parent
species which have contributed to the origin of the hybrids. Our
tuberous begonias owe their variability to at least seven original
parent species, [273] and to the almost incredible number of
combinations which are possible between their characters. The first of
these crosses was made in the nursery of Veitch and Sons near London by
Seden, and the first hybrid is accordingly known as _Begonia sedeni_ and
is still to be met with. It has been superseded by subsequent crosses
between the _sedeni_ itself and the _Veitchi_ and _rosiflora_, the
_davisii_, the _clarkii_ and others. Each of them contributed its
advantageous qualities, such as round flowers, rosy color, erect flower
stalks, elevation of the flowers above the foliage and others. New
crosses are being made continuously, partly between the already existing
hybrids and partly with newly introduced wild species. Only rarely is it
possible to get pure seeds, and I have not yet been able to ascertain
whether the hybrids would come true from seed. Specific and varietal
characters may occur together in many of the several forms, but nothing
is as yet accurately known as to their behavior in pure fertilizations.
Constancy and segregation are thrown together in such a manner that
extreme variability results, and numerous beautiful types may be had,
and others may be expected from further crosses. For a scientific
analysis, however, the large range of recorded facts and the written
history, which at first sight [274] seems to have no lacunae, are not
sufficient. Most of the questions remain open and need investigation. It
would be a capital idea to try to repeat the history of the begonias or
any other hybrid race, making all the described crosses and then
recording the results in a manner requisite for complete and careful
scientific investigations.
Many large genera of hybrid garden-flowers owe their origin to species
rich in varieties or in elementary subspecies. Such is the case with the
gladiolus and the tulips. In other cases the original types have not
been obtained from the wild state but from the cultures of other
countries.
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