In most cases of wild, and of garden-varieties, the relation between
them and the parent-species rests upon comparative evidence. Often the
variety is known to be younger, in other cases it may be only of local
occurrence, but ordinarily the historic facts about its origin have
never been known or have long since been forgotten.
The easiest and most widely known varietal crosses are those between
varieties with white flowers and the red- or blue-flowered species. Here
the color prevails in the hybrid over the lack of pigment, and as a rule
the hybrid is as deeply tinted as the species itself, and cannot be
distinguished from it, without an investigation of its hereditary
qualities. Instances may be cited of the white varieties of the
snapdragon, of the red clover, the long-spurred violet (_Viola_ [282]
_cornuta_) the sea-shore aster (_Aster Tripolium_), corn-rose
(_Agrostemma Githago_), the Sweet William (_Silene Armeria_), and many
garden flowers, as for instance, the _Clarkia pulchella_, the
_Polemonium coeruleum_, the _Veronica longifolia_, the gloxinias and
others. If the red hue is combined with a yellow ground-color in the
species, the variety will be yellow and the hybrid will have the red and
yellow mixture of the species as for instance, in the genus _Geum_. The
toad-flax has an orange-colored palate, and a variety occurs in which
the palate is of the same yellow tinge as the remaining parts of the
corolla. The hybrid between them is in all respects like the
parent-species.
Other instances could be given. In berries the same rule prevails. The
black nightshade has a variety with yellow berries, and the black color
returns in the hybrid. Even the foliage of some garden-plants may afford
instances, as for instance, the purplish amaranth (_Amaranthus
caudatus_). It has a green variety, but the hybrid between the two has
the red foliage of the species.
Special marks in leaves and in flowers follow the same rule. Some
varieties of the opium poppy have large black patches at the basal end
of the petals, while in others this pattern is entirely white. In
crossing two such varieties, [283] for instance, the dark "Mephisto"
with the white-hearted "Danebrog," the hybrid shows the active character
of the dark pattern.
Hairy species crossed with their smooth varieties produce hairy hybrids,
as in some wheats, in the campion (_Lychnis_), in _Biscutella_ and
others. The same holds good for the crosses between spiny species and
their unarmed derivatives, as in the thorn-apple, the corn-crowfoot
(_Ranunculus arvensis_) and others.
Lack of starch in seeds is observed in some varieties of corn and of
peas. When such derivatives are crossed with ordinary starch-producing
types, the starch prevails in the hybrid.
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