This simple law for the constitution of the second generation of
varietal hybrids with a single differentiating mark in their parents is
called the law of Mendel. Mendel published it in 1865, but his paper
remained nearly unknown to scientific hybridists. It is only of late
years that it has assumed a high place in scientific literature, and
attained the first rank as an investigation on fundamental questions of
heredity. [294] Read in the light of modern ideas on unit characters it
is now one of the most important works on heredity and has already
widespread and abiding influence on the philosophy of hybridism in
general.
But from its very nature and from the choice of the material made by
Mendel, it is restricted to balanced or varietal crosses. It assumes
pairs of characters and calls the active unit of the pair dominant, and
the latent recessive, without further investigations of the question of
latency. It was worked out by Mendel for a large group of varieties of
peas, but it holds good, with only apparent exceptions, for a wide range
of cases of crosses of varietal characters. Recently many instances have
been tested, and even in many cases third and later generations have
been counted, and whenever the evidence was complete enough to be
trusted, Mendel's prophecy has been found to be right.
According to this law of Mendel's the pairs of antagonistic characters
in the hybrid split up in their progeny, some individuals reverting to
the pure parental types, some crossing with each other anew, and so
giving rise to a new generation of hybrids. Mendel has given a very
suggestive and simple explanation of his formula. Putting this in the
terminology of to-day, and limiting it to the occurrence of only [295]
one differential unit in the parents, we may give it in the following
manner. In fertilization, the characters of both parents are not
uniformly mixed, but remain separated though most intimately combined in
the hybrid throughout life. They are so combined as to work together
nearly always, and to have nearly equal influence on all the processes
of the whole individual evolution. But when the time arrives to produce
progeny, or rather to produce the sexual cells through the combination
of which the offspring arises, the two parental characters leave each
other, and enter separately into the sexual cells. From this it may be
seen that one-half of the pollen-cells will have the quality of one
parent, and the other the quality of the other. And the same holds good
for [296] the egg-cells. Obviously the qualities lie latent in the
pollen and in the egg, but ready to be evolved after fertilization has
taken place.
Public-domain text, read in full here on John Shaqi.
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