The corresponding experiment could be made with plants of a pure
sugar-race by pollination with hybrid pollen. The spikes would show
exactly the same mixture as in the above case, but now this may be
considered as conclusive proof that half the pollen-grains represent the
quality of one parent and the other half the quality of the other.
Another corollary of Mendel's law is the following. In each generation
two groups return to purity, and one-half remains hybrid. These last
will repeat the same phenomenon of splitting in their progeny, and it is
easily seen that the same rule will hold good for all succeeding
generations. According to Mendel's principle, in each year there is a
new hybridization, differing in no respect from the first and original
one. If the hybrids only are propagated, each year will show one-fourth
of the offspring returning to the specific character, one-fourth
assuming the type of the variety and one-half remaining hybrid. I have
tested this with a hybrid between the ordinary nightshade with black
berries, and its variety, _Solanum nigrum chlorocarpum_, with pale
yellow fruits. Eight generations of the hybrids were cultivated, [299]
disregarding always the reverting offspring. At the end I counted the
progeny of the sixth and seventh generations and found figures for their
three groups of descendants, which exactly correspond to Mendel's
formula.
Until now we have limited ourselves to the consideration of single
differentiating units. This discussion gives a clear insight into the
fundamental phenomena of hybrid fertilization. It at once shows the
correctness of the assumption of unit-characters, and of their pairing
in the sexual combinations.
But Mendel's law is not at all restricted to these simple cases. Quite
on the contrary, it explains the most intricate questions of
hybridization, providing they do not transgress the limits of
symmetrical unions. But in this realm nearly all results may be
calculated beforehand, on the ground of the principle of probability.
Only one more assumption need be discussed. The several pairs of
antagonistic characters must be independent from, and uninfluenced by,
one another. This premise seems to hold good in the vast majority of
cases, though rare exceptions seem to be not entirely wanting. Hence the
necessity of taking all predictions from Mendel's law only as
probabilities, which will prove true in most, but not necessarily in all
cases. [300] But here we will limit ourselves to normal cases.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account