In order to look closer into the real mechanism of this form of
variability, and of this constant tendency to occasional reversions, it
will be best to limit ourselves first to a single case, and to try to
gather all the evidence, which can be obtained by an examination of the
hereditary relations of its sundry constituents.
[314] This may best be done by determining the degree of inheritance for
the various constituents of the race during a series of years. It is
only necessary to apply the two precautions of excluding all
cross-fertilization, and of gathering the seeds of each individual
separately. We do not need to ascertain whether the variety as such is
permanent; this is already clear from the simple fact of its antiquity
in so many cases. We wish to learn what part each individual, or each
group of individuals with similar characters, play in the common line of
inheritance. In other words, we must build up a genealogical tree,
embracing several generations and a complete set of the single cases
occurring within the variety, in order to allow of its being considered
as a part of the genealogy of the whole. It should convey to us an idea
of the hereditary relations during the life-time of the variety.
It is manifest that the construction of such a genealogical tree
requires a number of separate experiments. These should be extended over
a series of years. Each should include a number of individuals large
enough to allow the determination of the proportion of the different
types among the offspring of a single plant. A species which is easily
fertilized by its own pollen, and which bears capsules with [315] large
quantities of seeds, obviously affords the best opportunities. As such,
I have chosen the common snapdragon of the gardens, _Antirrhinum majus_.
It has many striped varieties, some tall, others of middle height, or of
dwarfed stature. In some the ground-color of the flowers is yellow, in
others it is white, the yellow disappearing, with the exception of a
large mark in the throat. On these ground-colors the red pigment is seen
lying in streaks of pure carmine, with white intervals where the yellow
fails, but combined with yellow to make a fiery red, and with yellow
intervals when that color is present. This yellow color is quite
constant and does not vary in any marked degree, notwithstanding the
fact that it seems to make narrower and broader stripes, according to
the parts of the corolla left free by the red pigment. But it is easily
seen that this appearance is only a fallacious one.
The variety of snapdragon chosen was of medium height and with the
yellow ground-color, and is known by horticulturists as _A. majus luteum
rubro-striatum_. As the yellow tinge showed itself to be invariable; I
may limit my description to the red stripes.
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