Similar cases occur not rarely among cultivated plants. In the wild
state they seem to be wholly wanting. Neither are they met with as
occasional anomalies nor as distinct varieties. On the contrary, many
garden-flowers that are colored in the species, and besides this have a
white or yellow variety, have also striped sorts. The oldest instance is
probably the marvel of Peru, _Mirabilis Jalappa_, which already had more
than one striped variety at the time of its introduction from Peru into
the European gardens, about the beginning of the seventeenth century.
Stocks, liver-leaf (_Hepatica_), dame's violet (_Hesperis_), Sweet
William (_Dianthus barbatus_), and periwinkles (_Vinca minor_) seem to
be in the same condition, as their striped varieties were already quoted
[323] by the writers of the same century. Tulips, hyacinths, _Cyclamen_,
_Azalea_, _Camellia_, and even such types of garden-plants as the meadow
crane's-bill (_Geranium pratensev) have striped varieties. It is always
the red or blue color which occurs in stripes, the underlying ground
being white or yellow, according to the presence or absence of the
yellow in the original color mixture.
All these varieties are known to be permanent, coming true during long
series of successive generations. But very little is known concerning
the more minute details of their hereditary qualities. They come from
seed, when this is taken from striped individuals, and thence revert
from time to time to the corresponding monochromatic type. But whether
they would do so when self-fertilized, and whether the reversionary
individuals are always bound to return towards the center of the group
or towards the opposite limit, remains to be investigated. Presumably
there is nowhere a real transgression of the limits, and never or only
very rarely and at long intervals of time a true production of another
race with other hereditary qualities.
In order to satisfy myself on these points, I made some
pedigree-cultures with the striped forms of dame's violet (_Hesperis
matronalis_) [324] and of _Clarkia pulchella_. Both of them are
ever-sporting varieties. The experiments were conducted during five
generations with the violet, and during four with the striped Clarkia,
including the progeny of the striped and of the monochromatic red
offspring of a primitive striped plant. I need not give the figures here
for the numerical relations between the different types of each group,
and shall limit myself to the statement that they behaved in exactly the
same manner as the snapdragon.
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