The change of stamens into petals is a gradual modification. All
intermediate steps are easily to be found. In some flowers all stamens
may be enlarged, in others only part of them. Often the broadened
filaments bear one or two fertile anthers. The fertility is no doubt
diminished, but not wholly destroyed. Individual specimens may occur,
which cannot produce any seed, but then others of the same lot may be as
fertile as can be desired. As a whole, such double varieties are
regularly propagated by seed.
Petalomany is the tendency of the axis of some flowers never to make any
stamens or pistils, not even in altered or rudimentary form. Instead of
these, they simply continue producing petals, going on with this
production without any other limit than the supply of available food.
Numerous petals fill the entire space within the outer rays, and in the
heart of the flower innumerable young ones are developed half-way, not
obtaining food enough to attain [331] full size. Absolute sterility is
the natural consequence of this state of things.
Hence it is impossible to have races of petalomanous types. If the
abnormality happens to show itself in a species, which normally
propagates itself in an asexual way, the type may become a vegetative
variety, and be multiplied by bulbs, buds or cuttings, etc. Some
cultivated anemones and crowfoots (_Ranunculus_) are of this character,
and even the marsh-marigold (_Caltha palustris_) has a petalomanous
variety. I once found in a meadow such a form of the meadow-buttercup
(_Ranunculus_ acris_), and succeeded in keeping it in my garden for
several years, but it did not make seeds and finally died. Camellias are
known to have both types of double flowers. The petalomanous type is
highly regular in structure, so much so as to be too uniform in all its
parts to be pleasing, while the conversion of stamens into petals in the
alternative varieties gives to these flowers a more lively diversity of
structure. Lilies have a variety called _Lilium candidum flore pleno_,
in which the flowers seem to be converted into a long spike of bright,
white narrow bracts, crowded on an axis which never seems to cease their
production.
It is manifestly impossible to decide how all such sterile double
flowers have originated. [332] Perhaps each of them originally had a
congruent single-flowered form, from which it was produced by seed in
the same way as the double stocks now are yearly. If this assumption is
right, the corresponding fertile line is now lost; it has perhaps died
out, or been masked. But it is not absolutely impossible that such
strains might one day be discovered for one or another of these now
sterile varieties.
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