Obviously it is not always easy to decide definitely whether a given
individual belongs to such a race or not. Many trials may be necessary
to secure the special race. I had the good fortune to find two plants of
clover, bearing one quinate and several quaternate leaves, on an
excursion in the neighborhood of Loosdrecht in Holland. After
transplanting them into my garden, I cultivated them during three years
and observed a slowly increasing number of anomalous leaves. This number
in one summer amounted to 46 quaternate and 16 quinate leaves, and it
was evident that I had secured an instance of the rare "five-leaved"
race which I am about to describe.
Before doing so it seems desirable to look somewhat closer into the
morphological features of the problem. Pinnate and palmate leaves often
vary in the number of their parts. This variability is generally of the
nature of a common fluctuation, the deviations grouping themselves
around an average type in the ordinary way. Ash leaves bear five pairs,
and [342] the mountain-ash (_Sorbus Aucuparia_) has six pairs of
leaflets in addition to the terminal one. But this number varies
slightly, the weaker leaves having less, the stronger more pairs than
the average. Such however, is not the case, with ternate leaves, which
seem to be quite constant. Four leaflets occur so very rarely that one
seems justified in regarding them rather as an anomaly than, as a
fluctuation. And this is confirmed by the almost universal absence of
two-bladed clover-leaves.
Considering the deviation as an anomaly, we may look into its nature.
Such an inquiry shows that the supernumerary leaflets owe their origin
to a splitting of one or more of the normal ones. This splitting is not
terminal, as is often the case with other species, and as it may be seen
sometimes in the clover. It is for the most part lateral. One of the
lateral nerves grows out becoming a median nerve of the new leaflet.
Intermediate steps are not wanting, though rare, and they show a gradual
separation of some lateral part of a leaflet, until this division
reaches the base and divides the leaflet into two almost equal parts. If
this splitting occurs in one leaflet we get the "four-leaved" Clover, if
it occurs in two there will be five leaflets. And if, besides this, the
terminal leaflet produces a derivative on one or both of its sides,
[343] we obtain a crown of six or seven leaflets on one stalk. Such were
often met with in the race I had under cultivation, but as a rule it did
not exceed this limit.
The same phenomenon of a lateral doubling of leaflets may of course be
met with in other instances. The common laburnum has a variety which
often produces quaternate and quinate leaves, and in strawberries I have
also seen instances of this abnormality. It occurs also in pinnate
leaves, and complete sets of all the intermediate links may often be
found on the false or bastard-acacia (_Robinia Pseud_Acacia_).
Public-domain text, read in full here on John Shaqi.
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