Returning to the original plants of this race, it is evidently
impossible to decide whether they were really the beginning of a new
strain, and had originated themselves by some sudden change from the
common type, or whether they belonged to an old variety, which had
propagated itself perhaps during centuries, unobserved by man. But the
same difficulty generally arises when new varieties are discovered. Even
the behavior of the plants themselves or of their progeny does not
afford any means of deciding the question. The simplest way of stating
the matter therefore, is to say that I accidentally found two
individuals of the "five-leaved" race. By transplanting them into my
garden, I have isolated them and kept them free from cross-fertilization
with the ordinary type. Moreover, I have brought them under such
conditions as are necessary for the full development of their
characters. And last but not least, I have tried to improve this
character as far as possible by a very rigid and careful selection.
The result of all this effort has been a rapid improvement of my strain.
I saved the seed of the original plants in 1889 and cultivated the
second generation in the following year. It [346] showed some increase
of the anomaly, but not to a very remarkable degree. In the flowering
period I selected four plants with the largest number of quaternate and
quinate leaves and destroyed all the others. I counted in the average 25
anomalous organs on each of them. From their seed I raised the third
generation of my culture in the year 1891.
This generation included some 300 plants, on which above 8,000 leaves
were counted. More than 1,000 were quaternate or quinate, the ternate
leaves being still in the majority. But the experiment clearly showed
that "four-leaved" clovers may be produced in any desired quantity,
provided that the seed of the variety is available. In the summer only
three, four and five leaflets on one stalk were seen, but towards the
fall, and after the selection of the best individuals, this number
increased and came up to six and seven in some rare instances.
The selection in this year was by no means easy. Nearly all the
individuals produced at least some quaternate leaves, and thereby showed
the variety to be quite pure. I counted the abnormal organs on a large
group of the best plants, and selected 20 excellent specimens from them,
with more than one-third of all their leaves changed in the desired
manner. Having brought my race up to this point, I [347] was able to
introduce a new and far more easy mark, afforded by the seedlings, for
my selections. This mark has since remained constant, and has brought
about a rapid continuance of the improvement, without necessitating such
large cultures.
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