I wish to point out that the decision of what is to be expected from
deviating specimens may become manifest within one or two generations.
Even the generation grown from the seeds of [361] the first observed
aberrant-individuals, if gathered after sufficient isolation during the
period of blossoming, may show which type of inheritance is present,
whether it is an unpromising half-race, or a richly endowed sporting
variety. I have kept such strains repeatedly after the first isolation,
and a special case, that of cotyledoneous aberrations, will be dealt
with later. The first generation always gave a final decision, provided
that a suitable method of cultivation for the species under observation
was found at the beginning. This however, is a condition, which it is
not at all easy to comply with, when new sorts are introduced into a
garden. Especially so when they had been collected in the wild state.
Often one or two years, sometimes more, are necessary to find the proper
method of sowing, manuring, transplanting and, other cultural methods
satisfactory to the plants. Many wild species require more care and more
manure in gardens than the finest garden flowers. And a large number are
known to be dependent on very particular conditions of soil.
One of the most curious features of anomalies, which has been learned
from accumulated instances, is the fact that they obey definite laws as
to their occurrence on the different parts of the plant. Obviously such
laws are [362] not apparent as long as each plant produces only one or
two, or, at most, a few instances of the same deviation. On the
contrary, any existing regularity must betray itself, as soon as a
larger number of instances is produced. A rule of periodicity becomes
most clearly manifest in such cases.
This rule is shown by no other race in a more undoubted and evident
manner than by the "five-leaved" clover. Evidently the several degrees
of deviation, going from three to seven leaflets, may be regarded as
responses to different degrees of variation, and their distribution over
the stems and branches, or over the whole plant, may be considered as
the manifestation of the ever-changing internal tendency to vary.
Considered from this point of view, my plants always showed a definite
periodicity in this distribution, which is the same for the whole plant.
Each of them, and each of the larger branches, begin with atavistic
leaves or with slight deviations. These are succeeded by greater
deviations, but only the strongest axes show as many as seven leaflets
on a stalk. This ordinarily does not occur before the height of
development is reached, and often only towards its close. Then the
deviation diminishes rapidly, returning often to atavistic leaves at the
summit of the stem or branch. I give the numbers of the [363] leaves of
a branch, in their order from the base to the top. They were as follows:
3. 4. 5. 6. 7. 5. 5. 4.
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