Peloric flowers are terminal in some cases, but occur in the lower parts
of the flower-spikes in others. Some varieties of gladiolus commence on
each spike with more or less double flowers, which, higher up, are
replaced by single ones. A wide range of bulbs and perennial
garden-plants develop their varietal characters only partly when grown
from seed and flowering for the first time. The annual
garden-forget-me-not of the Azores (_Myosotis azorica_) has a variety
with curiously enlarged flowers, often producing 20 or more
corolla-segments in one flower. But this number gradually diminishes as
the season advances. It would be quite superfluous to give further proof
of the general validity of the law of periodicity in ever-sporting
varieties.
[369]
LECTURE XIII
PISTILLODY IN POPPIES
One of the most curious anomalies that may be met with in ornamental
garden-plants is the conversion of stamens into pistils. It is neither
common nor rare, but in most cases the change is so slight comparatively
that it is ordinarily overlooked. In the opium-poppy, on the contrary,
it is very showy, and heightens the ornamental effect of the young
fruits after the fading of the flowers. Here the central capsule is
surrounded by a large crown of metamorphosed stamens.
This peculiarity has attracted the attention both of horticulturists and
of botanists. As a rule not all the stamens are changed in this way but
only those of the innermost rows. The outer stamens remain normal and
fertile, and the flowers, when pollinated with their own pollen, bear as
rich a harvest of seeds as other opium-poppies. The change affects both
the filament and the anther, the former of which is dilated into a
sheath. Within this sheath perfect [370] and more or less numerous
ovules may be produced. The anthers become rudimentary and in their
place broad leafy flaps are developed, which protrude laterally from the
tip and constitute the stigmas. Ordinarily these altered organs are
sterile, but in some instances a very small quantity of seed is
produced, and when testing their viability I succeeded in raising a few
plants from them.
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