The fact of inheritance is shown by the experience of many authors, and
by the circumstance already quoted, that the variety has been propagated
from seed for more than half a century, and may be obtained from various
seed merchants. In respect to the variability, the variety belongs to
the ever-sporting group, constituting a type which is more closely
related to the "five-leaved" clover than to the striped flowers or even
the double stocks.
It fluctuates around an average type with half filled crowns, going as
far as possible in both directions, but never transgressing either
limit. It is even doubtful whether the presumable limits are, under
ordinary circumstances, ever reached. Obviously one extreme would be the
conversion of all the stamens, and the other the absolute deficiency of
any marked tendency to such a change. Both may occur, and will probably
be met with from time to time. But they must be extremely rare, since in
my own extensive experiments, which were strictly controlled, I never
was able to find a single instance of either of them. Some of the outer
stamens have always remained unchanged, yielding enough pollen for the
artificial pollination of [375] the central ovary, and on the other hand
some rudiments of hardened filaments were always left, even if they were
reduced to small protuberances on the thalamus of the flower. Between
these extremes all grades occur. From single, partially or wholly
changed stamens upwards to 150 and over, all steps may be seen. It is a
true fluctuating variability. There is an average of between 50 and 100,
constituting a nearly filled crown around the central capsule. Around
this average the smaller deviations are most numerous and the larger
ones more rare. The inspection of any bed of the variety suffices to
show that, taken broadly, the ordinary laws of fluctuating variability
are applicable. No counting of the single individuals is required to
dispel all doubts on this point.
Moreover all intermediate steps respecting the conversion of the single
stamens may nearly always be seen. Rarely all are changed into normal
secondary ovaries with a stigma and with a cavity filled with ovules.
Often the stigma is incomplete or even almost wanting, in other
instances the ovules are lacking or the cavity itself is only partially
developed. Not rarely some stamens are reduced and converted into thin
hard stalks, without any appearance of an ovary at their tip. But then
the demarcation [376] between them and the thalamus fails, so that they
cannot be thrown off when the flower fades away, but remain as small
stumps around the base of the more fully converted filaments. This fact
would frequently render the enumeration of the altered organs quite
unreliable.
Public-domain text, read in full here on John Shaqi.
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